Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors: Neoteleostei
Infracohors: Eurypterygia
Sectio: Ctenosquamata
Subsectio: Acanthomorphata
Divisio/Superordo: Acanthopterygii
Subdivisio: Percomorphaceae
Series: Eupercaria
Ordo: Perciformes
Subordo: Percoidei
Superfamilia: Percoidea
Familia: Percidae
Subfamilia: Percinae
Genus: Gymnocephalus
Species: G. acerinus – G. ambriaelacus – G. baloni – G. cernua – G. schraetser
Name
Gymnocephalus Bloch, 1793
References
Gymnocephalus species list in FishBase,
Froese, R. & Pauly, D. (eds.) 2024. FishBase. World Wide Web electronic publication, www.fishbase.org, version 02/2024.
Taxon identifiers
Wikidata: Q927216 ADW: Gymnocephalus BOLD: 3692 CoL: 4RSB EoL: 24006 Fauna Europaea: 304850 Fauna Europaea (2): 99271b4d-0b18-497f-ae6f-d1429f4f68f0 GBIF: 2382125 iNaturalist: 88656 NBN: NHMSYS0000544679 NCBI: 39193 WoRMS: 151301
Vernacular names
беларуская (тарашкевіца): Джгіры
беларуская: Джгіры
čeština: ježdík
English: Gymnocephalus
français: Gymnocephalus
magyar: Gymnocephalus
italiano: Gymnocephalus
latviešu: Ķīšu ģints
Nederlands: Gymnocephalus
polski: Gymnocephalus
português: Gymnocephalus
русский: Ерши
svenska: Gärssläktet
українська: Йорж (рід)
Gymnocephalus is a genus of ray-finned fishes from the family Percidae, which includes the perches, pike-perches and darters. They are from the Western Palearctic area, although one species, Gymnocephalus cernua has been accidentally introduced to the Great Lakes region where it is regarded as an invasive species. They have the common name "ruffe" and resemble the true perches in the genus Perca, but are usually smaller and have a different pattern.
Characteristics
The species within the genus Gymnocephalus have a number of characters in common including that their dorsal fins are not completely separate, they have enlarged canals extending from the lateral line on their heads, the preorbital bone covers the maxillary bone, presence of setiform or bristle-like teeth, having very few or no vomerine and palatine teeth and the possession of three paired bones in the neck, known as extrascapulars, in their lateral line system, of which, two are simple tubes.[4]
Species
There are currently five recognized species in this genus:[5]
Gymnocephalus acerina (J. F. Gmelin, 1789) (Donets ruffe)
Gymnocephalus ambriaelacus Geiger & Schliewen, 2010
Gymnocephalus baloni Holčík & K. Hensel, 1974 (Balon's ruffe)
Gymnocephalus cernua (Linnaeus, 1758) (ruffe)
Gymnocephalus schraetser (Linnaeus, 1758) (schraetzer)
Taxonomy
Gymnocephalus was created by the German physician and naturalist Marcus Elieser Bloch (1723-1799) with Perca schraester as the type species. It has traditionally been placed in the subfamily Percinae alongside the true perch of the genus Perca.[5][2] However, Gymnocephalus appears to be the sister taxon to both the Percinae and to the Luciopercinae.[4] The 5th edition of Fishes of the World treats Gymnocephalus as the only genus in the monotypic subfamily Acerinae,[6] although Gill's Gymnocephalinae is referred to in some sources.[1] The name of the genus is a compound of the Greek gymno meaning "naked" and kephalos meaning "head".[7] Within the genus molecular studies have shown that the ruffe G. cernua is sister to a clade consisting of the Danube ruff G. baloni and the schraetzer or striped ruffe G. schraetser, and that these last two species originated from a common ancestor about 8 million years ago and it has also been suggested that the relatively newly described G. ambriaelacus may be synonymous with G. baloni. G. acerina has not had its genetics sampled which would assist understanding of the genus's phylogenetics.[4]
Geographic distribution
The species in the genus Gymnocephalus are found in Europe. One species, G. cernua, originating from the river Elbe in Germany, was accidentally introduced to North America in ballast water near the mouth of the St Louis River in Lake Superior.[4] This species has also been introduced outside of its native range in Europe.[8]
References
Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014). "Family-group names of Recent fishes". Zootaxa. 3882 (2): 001–230. doi:10.11646/zootaxa.3882.1.1. PMID 25543675.
Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Gymnocephalus". Catalog of Fishes. California Academy of Sciences. Retrieved 13 September 2020.
Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Genera in the family Percinae". Catalog of Fishes. California Academy of Sciences. Retrieved 13 September 2020.
Carol A. Stepien & Amanda Haponski (2015). "Taxonomy, Distribution, and Evolution of the Percidae". In Patrick Kestemont; Konrad Dabrowski & Robert C. Summerfelt (eds.). Biology and Culture of Percid Fishes. Springer, Dordrecht. pp. 3–60. doi:10.1007/978-94-017-7227-3_1. ISBN 978-94-017-7227-3.
Froese, Rainer and Pauly, Daniel, eds. (2014). Species of Gymnocephalus in FishBase. February 2014 version.
J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016). Fishes of the World (5th ed.). Wiley. pp. 448–450. ISBN 978-1-118-34233-6. Archived from the original on 2019-04-08. Retrieved 2020-09-13.
Froese, Rainer; Pauly, Daniel (eds.). "Gymnocephalus cernua year-2019". Fishbase.
Colin E. Adams; Peter S. Maitland (1998). "The Ruffe Population of Loch Lomond, Scotland: Its Introduction, Population Expansion, and Interaction with Native Species (abstract)". Journal of Great Lakes Research. 24 (2): 249–262. doi:10.1016/s0380-1330(98)70817-2.
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