Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Otomorpha
Subcohors: Ostariophysi
Sectio: Otophysa
Ordo: Cypriniformes
Subordo: Cyprinoidei
Familia: Acheilognathidae
Genus: Acheilognathus
Species: A. asmussii – A. barbatulus – A. barbatus – A. binidentatus – A. brevicaudatus – A. changtingensis – A. cyanostigma – A. deignani – A. elongatoides – A. elongatus – A. fasciodorsalis – A. gracilis – A. hondae – A. hypselonotus – A. imberbis – A. imfasciodorsalis – A. koreensis – A. kyphus – A. longibarbatus – A. longipinnis – A. macromandibularis – A. majusculus – A. melanogaster – A. meridianus – A. microphysa – A. nguyenvanhaoi – A. omeiensis – A. peihoensis – A. polylepis – A. polyspinus – A. rhombeus – A. signifer – A. somjinensis – A. striatus – A. tabira – A. taenianalis – A. tonkinensis – A. typus – A. yamatsutae
Name
Acheilognathus Bleeker, 1860: 427
Type species: Capoeta rhombea Temminck & Schlegel, 1846. Type by subsequent designation.
References
Bleeker, P. 1860: Conspectus systematis Cyprinorum. Natuurkundig Tijdschrift voor Nederlandsch Indië, v. 20: 421–441.
Naseka, A. M. and N. G. Bogutskaya 2004: Contribution to taxonomy and nomenclature of freshwater fishes of the Amur drainage area and the Far East (Pisces, Osteichthyes). Zoosystematica Rossica v. 12: 279–290, 2 pls.
Arai, R. , H. Fujikawa, and Y. Nagata 2007: Four new subspecies of Acheilognathus Bitterlings (Cyprinidae: Acheilognathidae) from Japan. Bulletin of the National Museum of Nature and Science (Ser. A) Suppl. 1: 1–28.
Saitoh, K., Shindo, K., Fujimoto, Y., Takahashi, K. & Shimada, T. 2016. Mitochondrial genotyping of an endangered bitterling Acheilognathus typus (Cyprinidae). ZooKeys 623: 131-141. DOI: 10.3897/zookeys.623.8981. Reference page.
Yang, Q.; Xiong, B.; Tang, Q.; Liu, H. 2010: Acheilognathus striatus (Family: Cyprinidae), a new bitterling species from the lower Yangtze River. Environmental biology of fishes, 88: 333–341. DOI: 10.1007/s10641-010-9645-6
Yang, Q.; Zhu, Y.; Xiong, B.; Liu, H. 2011: Acheilognathus changtingensis sp. nov., a new species of the cyprinid genus Acheilognathus (Teleostei: Cyprinidae) from southeastern China based on morphological and molecular evidence. Zoological science, 28(2): 158–167. DOI: 10.2108/zsj.28.158
Acheilognathus is a genus of cyprinid fish native to Asia. The name is derived from the Greek a, meaning "without", the Greek cheilos, meaning "lip", and the Greek gnathos, meaning "jaw".
Morphology
Fish of this genus range in length from 5 to 27 cm. They are similar in appearance to fishes of the genus Puntius.
Species
There are currently 41 recognized species in this genus:[1]
Acheilognathus asmussii (Dybowski, 1872) (Russian bitterling)
Acheilognathus barbatulus Günther, 1873
Acheilognathus barbatus Nichols, 1926
Acheilognathus brevicaudatus Y. R. Chen & Z. Y. Li, 1987
Acheilognathus changtingensis Qing Yang, Y. R. Zhu, B. X. Xiong & H. Z. Liu, 2011
Acheilognathus coreanus Steindachner, 1892
Acheilognathus cyanostigma D. S. Jordan & Fowler, 1903 (striped bitterling)
Acheilognathus deignani (H. M. Smith, 1945)
Acheilognathus elongatoides Kottelat, 2001
Acheilognathus elongatus (Regan, 1908) (elongate bitterling)
Acheilognathus fasciodorsalis V. H. Nguyễn, 2001
Acheilognathus gracilis Nichols, 1926
†Acheilognathus hondae (D. S. Jordan & Metz, 1913)
Acheilognathus hypselonotus (Bleeker, 1871)
Acheilognathus imberbis Günther, 1868
Acheilognathus imfasciodorsalis V. H. Nguyễn, 2001
Acheilognathus kyphus (Đ. Y. Mai, 1978)
Acheilognathus longibarbatus (Đ. Y. Mai, 1978)
Acheilognathus longipinnis Regan, 1905 (deepbody bitterling)
Acheilognathus macromandibularis A. Doi, R. Arai & H. Z. Liu, 1999
Acheilognathus macropterus (Bleeker, 1871)
Acheilognathus majusculus I. S. Kim & H. Yang, 1998
Acheilognathus melanogaster Bleeker, 1860 (Japanese bitterling)
Acheilognathus meridianus (H. W. Wu, 1939)
Acheilognathus microphysa J. X. Yang, X. L. Chu & Y. R. Chen, 1990
Acheilognathus nguyenvanhaoi H. D. Nguyễn, Đ. H. Trần & T. T. Tạ, 2013[2]
Acheilognathus omeiensis (H. J. Shih & T. L. Tchang, 1934)
Acheilognathus peihoensis Fowler, 1910
Acheilognathus polylepis (H. W. Wu, 1964)
Acheilognathus polyspinus (Holčík, 1972)
Acheilognathus rhombeus (Temminck & Schlegel, 1846)
Acheilognathus striatus Qing Yang, B. X. Xiong, Q. Y. Tang & H. Z. Liu, 2010
Acheilognathus tabira D. S. Jordan & W. F. Thompson, 1914
Acheilognathus tabira erythropterus R. Arai, Fujikawa & Nagata, 2007
Acheilognathus tabira jordani R. Arai, Fujikawa & Nagata, 2007
Acheilognathus tabira nakamurae R. Arai, Fujikawa & Nagata, 2007
Acheilognathus tabira tabira D. S. Jordan & W. F. Thompson, 1914
Acheilognathus tabira tohokuensis R. Arai, Fujikawa & Nagata, 2007
Acheilognathus taenianalis (Günther, 1873)
Acheilognathus tonkinensis (Vaillant, 1892)
Acheilognathus typus Bleeker, 1863
Acheilognathus yamatsutae T. Mori, 1928
In addition, there are two undescribed species in Korea awaiting further study:
Acheilognathus sp.[3]
Acheilognathus sp. HR[4]
References
Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Acheilognathus". FishBase. February 2013 version.
Nguyen Huu Duc; Tran Duc Hau; Ta Thi Thuy (2013). "A new species of genus Acheilognathus, Bleeker, 1895 from the Tien Yen river, Vietnam". Vietnam Journal of Biology. 35 (1): 18–22. doi:10.15625/0866-7160/v35n1.2933.
Chae, B.S.; Kim, S.K.; Lee, J.H.; Hwang, U.W. (2014). "Taxonomic status of Acheilognathus sp. (Cyprinidae) found in the Geum River, Korea". Korean Journal of Ichthyology (in Korean). 26 (4): 249–258. Retrieved 5 June 2023.
Kim, Y.H.; Yun, B.H.; Sung, M.S.; Bang, I.-C. (2022). "Taxonomic Status of Acheilognathus sp. HR (Cypriniformes: Acheilognathidae) Found in the Dalcheon River, a Tributary of Hangang River, Korea". Korean Journal of Ichthyology (in Korean). 34 (4): 231–243. doi:10.35399/ISK.34.4.2. Retrieved 5 June 2023.
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