Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Cladus: Theriimorpha
Cladus: Theriiformes
Cladus: Trechnotheria
Cladus: Zatheria
Subclassis: Theria
Cladus: Eutheria
Infraclassis: Placentalia
Magnordo: Boreoeutheria
Superordo: Laurasiatheria
Cladus: †Meridiungulata
Ordo: †Notoungulata
Genus: Isotemnidae
Genus: Pleurostylodon
Species: †P. modicus – †P. similis
Name
Pleurostylodon Ameghino 1897
References
Primary referencesAmeghino, F. (1897). Mammifères crétacés de l'Argentine: deuxième contribution à la connaisance de la faune mammalogique des couches à pyrotherium. Imprenta y papeleria "La Buenos Aires" Morena Esquina Peru.
Pleurostylodon is an extinct genus of notoungulate belonging to the family Isotemnidae. It lived during the Middle Eocene, in what is now Argentina.
Description
This genus is known from numerous remains, mainly cranial, allowing to reconstruct its morphology. It was approximately the size of a dog, with an appearance evocating a tapir or a boar.
Pleurostylodon had a large skull, widening in the orbital arch area, and narrowing in the posterior area of the muzzle, whose terminal part was enlarged and had small incisors ; there was no diastema after the canines. The muzzle was shorter than in more derived and specialized toxodonts such as Adinotherium, and the occipital area was narrower. Several characteristics of its maxilla, of its teeth and of its unspecialized auditory region evocates Homalodotherium.
The third upper incisor was enlarged, and vaguely resembling a canine, while the canine was larger and lanceolate. The premolars and molars had an external edge outside the sinuous protoloph, with a strong parastyle and a fold of the paracon. The protoloph and metaloph were complete, except in the first upper premolar, while the protocon and the hypocon were separated in the non-worn teeth. A sort of hook and several small crests were present in variable numbers in the median valley of the molars.
Classification
The genus Pleurostylodon was first described in 1897 by Florentino Ameghino, based on fossil remains found in Argentina in Middle Eocene terrains. The type species is Pleurostylodon modicus, but Ameghino described numerous other species in subsequent years, including P. complanatus, P. crassiramis, P. recticrista, P. similis.
Pleurostylodon is one of the better known genera of the family Isotemnidae, a group of basal notoungulates comprising numerous relatively unspecialized genera of variable size. The Isotemnidae family is considered by some to be paraphyletic, in which case Pleurostylodon may be closer to the basis of Toxodontia, which includes various derived genera such as Toxodon.
Bibliography
F. Ameghino. 1897. Mammiféres crétacés de l'Argentine (Deuxième contribution à la connaissance de la fauna mammalogique de couches à Pyrotherium) [Cretaceous mammals of Argentina (second contribution to the knowledge of the mammalian fauna of the Pyrotherium Beds)]. Boletin Instituto Geografico Argentino 18(4–9):406-521
F. Ameghino. 1901. Notices préliminaires sur des ongulés nouveaux des terrains crétacés de Patagonie [Preliminary notes on new ungulates from the Cretaceous terrains of Patagonia]. Boletin de la Academia Nacional de Ciencias de Córdoba 16:349-429
F. Ameghino. 1902. Notices préliminaires sur des mammifères nouveaux des terrains Crétacé de Patagonie {preliminary notes on new mammals from the Cretaceous terrains of Patagonia]. Boletin de la Academia Nacional de Ciencias de Córdoba 17:5-70
F. Ameghino. 1904. Nuevas especies de mamíferos, cretáceos y terciarios de la República Argentina [New species of mammals, Cretaceous and Tertiarty, from the Argentine Republic]. Anales de la Sociedad Cientifica Argentina 56–58:1-142
G. G. Simpson. 1967. The beginning of the age of mammals in South America. Part II. Bulletin of the American Museum of Natural History 137:1-260
G. Billet. 2011. Phylogeny of the Notoungulata (Mammalia) based on cranial and dental characters. Journal of Systematic Palaeontology. 9 (4): 481–97. doi:10.1080/14772019.2010.528456. OCLC 740994816.
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