Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cladus: Metatheria
Cladus: Marsupialiformes
Cohors: Marsupialia
Ordo: Didelphimorphia
Familia: Didelphidae
Subfamilia: Didelphinae
Tribus: Thylamyini
Genus: Chacodelphys
Species: Chacodelphys formosa
Name
Chacodelphys formosa (Shamel, 1930)
Holotype: USNM 236330, adult ♂, skin and skull.
Type locality: “Linda Vista near the Riacho Pilaga, about 200 km northwest of Formosa, Provincia Formosa, Argentina”.
Synonyms
Marmosa muscula Shamel, 1930: 83 [original description, preoccupied by Didelphys (Marmosa) musculus Cabanis, 1848]
Marmosa formosa Shamel, 1930: 311 [replacement name]
Chacodelphys formosa: Voss, Gardner & Jansa, 2004: 2 [name combination]
References
Primary references
Shamel, H.H. 1930. A new murine opossum from Argentina. Journal of the Washington Academy of Sciences 20: 83–84. BHL
Shamel, H.H. 1930. A new name for Marmosa muscula Shamel. Journal of Mammalogy 11(3): 311. DOI: 10.2307/1374153
Voss, R.S., Gardner, A.L. & Jansa, S.A. 2004. On the relationships of Marmosa formosa Shamel, 1930 (Marsupialia, Didelphidae), a phylogenetic puzzle from the Chaco of northern Argentina. American Museum Novitates 3442: 1–18. DOI: 10.1206/0003-0082(2004)442<0001:OTROMF>2.0.CO;2 Open access; hdl: 2246/2807 Open access Reference page.
Additional references
Teta, P. & Pardiñas, U.F.J. 2007. Mammalia, Didelphimorphia, Didelphidae, Chacodelphys formosa (Shamel, 1930): Range extension. Check List 3(4): 333–335. Full article (PDF)
Links
Chacodelphys formosa in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.
IUCN: Chacodelphys formosa (Vulnerable)
Vernacular names
English: Pygmy Opossum
español: Marmosa pigmea de El Chaco
The Chacoan pygmy opossum (Chacodelphys formosa) is a recently described genus and species of didelphimorph marsupial. The only species in Chacodelphys, C. formosa, was known until 2004 from only one specimen collected in 1920 in the Chaco of Formosa Province, Argentina.[1] The species is gaining popularity as a pocket pet.
Description
The Chacoan pygmy opossum is the smallest known species of didelphid. It has a head-body length of 68 mm, a tail of 55 mm and a hind foot of 11. It differs from the other "marmosine" genera (Marmosa, Monodelphis, Thylamys, Tlacuatzin, Gracilinanus, Marmosops, Lestodelphys) in having a long third manual digit, no distinctly tricolored pelage, a long fourth pedal digit, and a tail shorter than head-body. No other marmosine genera has this combination of characters. It is endemic to a small region of northern Argentina.
Taxonomic history
C. formosa was originally described as Marmosa muscula Shamel (1930a); however, this name is preoccupied, so Shamel (1930b) renamed it M. formosa. Afterwards, George Tate (1933) considered it a valid member of his "Elegans group" (=Thylamys) of Marmosa, whereafter it has been variously synonymized or treated as a distinct species of Marmosa or Thylamys until 1989, when Gardner & Creighton (1989) listed it as a synonym of Gracilinanus agilis, and then later separated from this species as G. formosus.[2] Finally, Voss et al. (2004) erected the new genus Chacodelphys for the species.
References
Teta, P.; de la Sancha, N.; Flores, D. (2016). "Chacodelphys formosa". IUCN Red List of Threatened Species. 2016: e.T136547A22175485. doi:10.2305/IUCN.UK.2016-3.RLTS.T136547A22175485.en. Retrieved 19 November 2021.
Gardner, A.L. (2005). "Order Didelphimorphia". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 7. ISBN 978-0-8018-8221-0. OCLC 62265494.
Gardner, A.L. & Creighton, G.K. 1989. A new generic name for Tate's microtarsus group of South American mouse opossums (Marsupialia: Didelphidae). Proceedings of the Biological Society of Washington 102:3–7.
Shamel, H.H. 1930a. A new murine opossum from Argentina. Journal of the Washington Academy of Sciences 20:83-84.
Shamel, H.H. 1930b. A new name for Marmosa muscula Shamel. Journal of Mammalogy 11:311.
Tate, G.H.H. 1933. A systematic revision of the marsupial genus Marmosa with a discussion of the adaptive radiation of the murine opossums (Marmosa). Bulletin of the American Museum of Natural History 66:1–250.
Voss, R.S., Gardner, A.L. & Jansa, S.A. 2004. On the relationships of "Marmosa" formosa Shamel 1930 (Marsupialia, Didelphidae), a phylogenetic puzzle from the chaco of northern Argentina. American Museum Novitates 3442:1-18, 2 June 2004.
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