Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Superordo: † Pterosauromorpha
Ordo: †Pterosauria
Subordo: †Pterodactyloidea
Superfamilia: Ornithocheiroidea
Familia: Incertae sedis
Genus: †Piksi
Species: Piksi barbarulna
Name
Piksi barbarulna Varricchio, 2002
Piksi is a potential pterosaur[1] genus containing the single species Piksi barbarulna (meaning "strange elbowed big bird", from Blackfoot piksi, "big bird" or, specifically, "chicken" and Latin barbarus "strange, outlandish" + ulna, elbow[2]). It lived roughly 75 million years ago in what is now Montana, United States. Known from parts of a right wing – the humerus, ulna and radius bones – the only specimens found so far are housed in the Museum of the Rockies (collection number MOR 1113). The genus Piksi is monotypic at present.
The fossils were found in 1991 by Gloria Jean Siebrecht in the Blackfeet Indian Reservation, namely at Bob's Vacation Site locality TM-088, Glacier County. Recovered from an old stratum of the upper Two Medicine Formation, they are probably from an individual that died in or near a small pool. It was described in 2002 by David J. Varricchio.
Description and classification
The bones are fragmentary and represent roughly the elbow area. Comparing the fossils' size to the wing bones of other ground birds, P. barbarulna would have been relatively small, with a wingspan reaching 1 m (3.3 ft).[1]
The original description of the fossils found its affinities unresolvable except that it was probably an ornithothoracine bird. Agnolin and Varricchio (2012) reinterpreted Piksi barbarulna as a pterosaur rather than a bird, most likely a member of Ornithocheiroidea.[3] However, it has since been noted that the humerus of Piksi possesses features which found in some theropods but not in any pterodactyloid pterosaurs.[4]
However, more recent studies have recovered it as a pterosaur again.[1]
Ecology
The deposit in which the bones were found was a silty claystone. This was formed from sediments deposited during what seems to have been a rather cool phase of the Late Cretaceous:[5] sea levels of the Western Interior Seaway at least were apparently very low for Mesozoic standards, though this may also have been due to strong tectonic uplift in the Cordilleran Overthrust Belt. The location was inland, with the Western Interior Seaway's coast at least 350 km (220 miles) away.
Judging from the stratigraphical, sedimentological and faunal data, the environment was thus probably semi-humid, possibly (seasonally?) semi-arid grassland or shrubland in a tropical or subtropical climate. The claystone apparently formed from sediment of a small floodplain, such as an ephemeral pool.[citation needed]
A diverse fauna utilized the location as a habitat. Other theropods were plentiful, including Troodon (the nest of which was recovered), tyrannosaurids, and dromaeosaurids. Orodromeus was also abundant; herds might have come to the pool to drink or breed, as adults, juveniles and hatchlings were found together. Early mammals – metatherians and multituberculates – occurred in the area, as well as lizards. That there was a temporary though not permanent body of water is indicated by the presence of articulated frog skeletons and the absence of fish and other aquatic animals.[citation needed]
See also
List of pterosaur genera
Timeline of pterosaur research
References
Longrich, N.R., Martill, D.M., and Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology, 16(3): e2001663. doi:10.1371/journal.pbio.2001663
(Varricchio 2002)
Federico L. Agnolin and David Varricchio (2012). "Systematic reinterpretation of Piksi barbarulna Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird" (PDF). Geodiversitas. 34 (4): 883–894. doi:10.5252/g2012n4a10. S2CID 56002643.
Martin-Silverstone, E.; Witton, M.P.; Arbour, V.M.; Currie, P.J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8): 160333. Bibcode:2016RSOS....360333M. doi:10.1098/rsos.160333. PMC 5108964. PMID 27853614.
Though still much warmer than today: see Cretaceous for contemporary climate.
Sources
Senter, Phil (2006): Scapular orientation in theropods and basal birds, and the origin of flapping flight. Acta Palaeontologica Polonica 51(2): 305–313. PDF fulltext
Snow, David W.; Perrins, Christopher M.; Doherty, Paul & Cramp, Stanley (1998): The complete birds of the western Palaearctic on CD-ROM. Oxford University Press. ISBN 0-19-268579-1
Varricchio, David J (2002). "A new bird from the Upper Cretaceous Two Medicine Formation of Montana". Canadian Journal of Earth Sciences. 39 (1): 19–26. doi:10.1139/e01-057.
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