Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Galliformes
Familia: Phasianidae
Subfamilia: Meleagridinae - Perdicinae - Phasianinae - Tetraoninae
Genera: Afropavo – Alectoris – Ammoperdix – Arborophila – Argusianus – Bambusicola – Bonasa – Caloperdix – Campocolinus – Canachites – Catreus – Centrocercus – Chrysolophus – Coturnix – Crossoptilon – Dendragapus – Falcipennis – Francolinus – Galloperdix – Gallus – Haematortyx – Ithaginis – Lagopus – Lerwa – Lophophorus – Lophura – Lyrurus – Margaroperdix – Melanoperdix – Meleagris – †Ophrysia – Ortygornis – Pavo – Peliperdix – Perdicula – Perdix – Phasianus – Polyplectron – Pternistis – Pucrasia – Rheinardia – Rhizothera – Rollulus – Scleroptila – Synoicus – Syrmaticus – Tetrao – Tetraogallus – Tetraophasis – Tetrastes – Tragopan – Tropicoperdix – Tympanuchus – Xenoperdix
Genera synonymized: Anurophasis – Excalfactoria – into Synoicus; Dendroperdix – into Ortygornis
Name
Phasianidae Horsfield, 1821
Typus: Phasianus Linnaeus, 1758
References
Primary references
Horsfield, T. 1821. Systematic Arrangement and Description of Birds from the Island of Java. Transactions of the Linnean Society of London (1)13: 133–200. BHL DOI: 10.1111/j.1095-8339.1821.tb00061.x Reference page. p. 185
Additional references
Seabrook-Davison, M., Huynen, L., Lambert, D.M. & Brunton, D.H. 2009. Ancient DNA resolves identity and phylogeny of New Zealand’s extinct and living quail (Coturnix sp.). PLOS ONE 4(7): e6400. DOI: 10.1371/journal.pone.0006400 Open access Reference page.
Kimball, R.T., St. Mary, C.M. & Braun, E.L. 2011. A Macroevolutionary Perspective on Multiple Sexual Traits in the Phasianidae (Galliformes). International Journal of Evolutionary Biology 11: 423938. DOI: 10.4061/2011/423938 Open access Reference page.
Kan, X.-Z., et al. 2010. Phylogeny of major lineages of galliform birds (Aves: Galliformes) based on complete mitochondrial genomes. Genet. Mol. Res. 9 (3): 1625-1633 DOI: 10.4238/vol9-3gmr898
Shen, Y.Y., et al. 2010. A mitogenomic perspective on the ancient, rapid radiation in the Galliformes with an emphasis on the Phasianidae. BMC Evolutionary Biology 10: 132 DOI: 10.1186/1471-2148-10-132
Hosner, P.A., et al. 2016. Avoiding Missing Data Biases in Phylogenomic Inference: An Empirical Study in the Landfowl (Aves: Galliformes). Mol. Biol. Evol. 33 (4): 1110–1125 DOI: 10.1093/molbev/msv347.
Chen, D., Liu, Y., Davison, G.W.H., Dong, L., Chang, J., Gao, S., Li, S-H. & Zhang, Z-W. 2015. Revival of the genus Tropicoperdix Blyth 1859 (Phasianidae, Aves) using multilocus sequence data. Zoological Journal of the Linnean Society 1175(2): 429–438 DOI: 10.1111/zoj.12273 Open access Reference page.
Hosner, P.A., Tobias, J.A., Braun, E.L. & Kimball, R.T. 2017. How do seemingly non-vagile clades accomplish trans-marine dispersal? Trait and dispersal evolution in the landfowl (Aves: Galliformes). Proceedings of the Royal Society B: Biological Sciences. 284 (1854): 20170210 DOI: 10.1098/rspb.2017.0210 Open access Reference page.
Chen, D., Braun, E.L., Forthman, M., Kimball, R.T. & Zhang, Z-W. 2018. A simple strategy for recovering ultraconserved elements, exons, and introns from low coverage shotgun sequencing of museum specimens: Placement of the partridge genus Tropicoperdix within the Galliformes. Molecular Phylogenetics and Evolution 129: 304–314 DOI: 10.1016/j.ympev.2018.09.005 Paywall Reference page.
Mandiwana-Neudani, T.G., Little, R.M., Crowe, T.M. & Bowie, R.C.K. 2019b. Taxonomy, phylogeny and biogeography of African spurfowls Galliformes, Phasianidae, Phasianinae, Coturnicini: Pternistis spp. Ostrich – Journal of African Ornithology 90(2): 145–172. DOI: 10.2989/00306525.2019.1584925 ResearchGate Reference page.
Mandiwana-Neudani, T.G., Little, R.M., Crowe, T.M. & Bowie, R.C.K. 2019b. Taxonomy, phylogeny and biogeography of ‘true’ francolins: Galliformes, Phasianidae, Phasianinae, Gallini; Francolinus, Ortygornis, Afrocolinus gen. nov., Peliperdix and Scleroptila spp. Ostrich – Journal of African Ornithology 90(3): 191–221. DOI: 10.2989/00306525.2019.1632954 ResearchGate Reference page.
Kimball, R.T., Hosner, P.A. & Braun, E.L. 2021. A phylogenomic supermatrix of Galliformes (Landfowl) reveals biased branch lengths. Molecular Phylogenetics and Evolution. 158: 107091. DOI: 10.1016/j.ympev.2021.107091 Paywall Reference page.
Vernacular names
беларуская: Фазанавыя
Deutsch: Fasanenartige
Esperanto: Fazanedoj
suomi: Aitokanat
Nordfriisk: Fasaanen
hrvatski: Fazanke
日本語: キジ科
lietuvių: Fazaniniai
македонски: Фазани
Nederlands: Fazantachtigen
norsk: Fasanfamilien
polski: Kurowate
русский: Фазановые
svenska: Fälthöns
ไทย: ไก่ฟ้า
Türkçe: Sülüngiller
українська: Фазанові
walon: Famile ås faizans
中文: 雉科
The Phasianidae are a family of heavy, ground-living birds, which includes pheasants, partridges, junglefowl, chickens, turkeys, Old World quail, and peafowl. The family includes many of the most popular gamebirds.[1] The family includes 185 species divided into 54 genera. It was formerly broken up into two subfamilies, the Phasianinae and the Perdicinae. However, this treatment is now known to be paraphyletic and polyphyletic, respectively, and more recent evidence supports breaking it up into two subfamilies: Rollulinae and Phasianinae, with the latter containing multiple tribes within two clades. The New World quail (Odontophoridae) and guineafowl (Numididae) were formerly sometimes included in this family, but are now typically placed in families of their own; conversely, grouse and turkeys, formerly often treated as distinct families (Tetraonidae and Meleagrididae, respectively), are now known to be deeply nested within Phasianidae, so they are now included in the present family.
Description
Phasianids are terrestrial. They range in weight from 43 g (1.5 oz) in the case of the king quail to 6 kg (13 lb) in the case of the Indian peafowl. If turkeys are included, rather than classified as a separate family, then the considerably heavier wild turkey capably reaches a maximum weight of more than 17 kg (37 lb). Length in this taxonomic family can vary from 12.5 cm (4.9 in) in the king quail up to 300 cm (120 in) (including the elongated train) in green peafowl, thus they beat even the true parrots in length diversity within a family of birds.[1][2] Generally, sexual dimorphism is greater in larger-sized birds, with males tending to be larger than females. They are generally plump, with broad, relatively short wings and powerful legs. Many have a spur on each leg, most prominently with junglefowl (including chickens), pheasants, turkeys, and peafowl. Some, like quails, partridges, and grouse, have reduced spurs to none at all. A few have two spurs on each of their legs instead of one, including peacock-pheasants and spurfowl. The bill is short and compact, particularly in species that dig deep in the earth for food such as the Mearns quail. Males of the bigger galliform species often boast brightly-coloured plumage, as well as facial ornaments such as combs, wattles, and/or crests.[citation needed]
Distribution and habitat
The Phasianidae are mostly an Old World family, with a distribution that includes most of Europe and Asia (except the far north), all of Africa except the driest deserts, and south into much of eastern Australia and (formerly) New Zealand. The Meleagridini (turkeys) are native to the New World, while the Tetraonini (grouse) are circumpolar; both of these are members of Phasianinae. The greatest diversity of species is in Southeast Asia and Africa. The Congo peacock is specific to the African Congo.
Overall, Rollulinae is restricted to the tropics of East and Southeast Asia and the mountains of Tanzania, Phasianinae have a circumpolar range in the temperate zones of both Eurasia and North America (but also range into the tropics of east and southeast Asia), and Pavoninae have a wide range across Africa, Eurasia, and Australasia in both temperate and tropical zones.
The family is generally sedentary and resident, although some members of the group undertake long migrations, like ptarmigans and Old World quail. Several species in the family have been widely introduced around the world, particularly pheasants, which have been introduced to Europe, Australia, and the Americas, specifically for hunting purposes. Captive populations of peafowl, domestic chickens, and turkeys have also escaped or been released and became feral.
Behaviour and ecology
The phasianids have a varied diet, with foods taken ranging from purely vegetarian diets of seeds, leaves, fruits, tubers, and roots, to small animals including insects, insect grubs, and even small reptiles. Most species either specialise in feeding on plant matter or are predatory, although the chicks of most species are insectivorous.
In addition to the variation in diet, a considerable amount of variation exists in breeding strategies among the Phasianidae. Compared to birds in general, a large number of species do not engage in monogamy (the typical breeding system of most birds). The francolins of Africa and some partridges are reportedly monogamous, but polygamy has been reported in the pheasants and junglefowl, some quail, and the breeding displays of peacocks have been compared to those of a lek. Nesting usually occurs on the ground; only the tragopans nest higher up in trees or stumps of bushes. Nests can vary from mounds of vegetation to slight scrapes in the ground. As many as 20 eggs can be laid in the nest, although 7-12 are the more usual numbers, with smaller numbers in tropical species. Incubation times can range from 14–30 days depending on the species, and is almost always done solely by the hen, although a few involve the male partaking in caring for the eggs and chicks, like the willow ptarmigan and bobwhite quail.
Relationship with humans
The red junglefowl of Southeast Asia is the wild ancestor of the domesticated chicken, the most important bird in agriculture, and the wild turkey similarly is the ancestor of the domestic turkey. Several species of pheasants and partridges are extremely important to humans. Ring-necked pheasants, several partridge and quail species, and some francolins have been widely introduced and managed as game birds for hunting. Several species are threatened by human activities.
Systematics and evolution
See also: List of Galliformes
The clade Phasianidae is the largest of the branch Galliformes, comprising 185 species divided into 54 genera.[3] This group includes the pheasants and partridges, junglefowl chickens, quail, and peafowl. Turkeys and grouse have also been recognized as having their origins in the pheasant- and partridge-like birds.
Until the early 1990s, this family was broken up into two subfamilies: the Phasianinae, including pheasants, tragopans, junglefowls, and peafowls;[4] and the Perdicinae, including partridges, Old World quails, and francolins.[5] Molecular phylogenies have shown that these two subfamilies are not each monophyletic, but actually constitute only one lineage with one common ancestor.[6][7] For example, some partridges (genus Perdix) are more closely affiliated to pheasants, whereas Old World quails and partridges from the genus Alectoris are closer to junglefowls.[6][7]
The earliest fossil records of phasianids date to the late Oligocene epoch, about 30 million years ago.[8]
Recent genera
Taxonomy and ordering is based on Kimball et al., 2021, which was accepted by the International Ornithological Congress. Tribes and subfamily names are based on the 4th edition of the Howard and Moore Complete Checklist of the Birds of the World. Genera without a tribe are considered to belong to tribe incertae sedis.[6][9][10][11]
Subfamily Rollulinae
Xenoperdix Dinesen et al., 1994 (forest partridges)
Caloperdix Blyth, 1861 (ferruginous partridge)
Rollulus Bonnaterre, 1791 (crested partridges)
Melanoperdix Jerdon, 1864 (black partridge)
Arborophila Hodgson, 1837 (hill partridges)
Subfamily Phasianinae
Phasianinae "Erectile clade"
Lerwa Hodgson, 1837 (snow partridge)
Ithaginis Wagler, 1832 (blood pheasant)
Tribe Lophophorini
Tragopan Cuvier, 1829 non Gray 1841 (tragopans)
Tetraophasis Elliot, 1871 (monal-partridges)
Lophophorus Temminck, 1813 non Agassiz 1846 (monals)
Pucrasia Gray, 1841 (koklass pheasant)
Tribe Tetraonini
Meleagris Linnaeus, 1758 (turkeys)
Bonasa Stephens, 1819 (ruffed grouse)
Tetrastes Keyserling & Blasius, 1840 (hazel grouse)
Centrocercus Swainson, 1832 (sage-grouse)
Dendragapus Elliot, 1864 (blue grouse)
Tympanuchus Gloger, 1841 (prairie-chickens and sharp-tailed grouse)
Lagopus Brisson, 1760 (ptarmigans)
Falcipennis Elliot, 1864 (Siberian grouse)
Canachites Stejneger, 1885 (spruce grouse)
Tetrao Linnaeus, 1758 (capercaillies)
Lyrurus Swainson, 1832 (black grouse)
Rhizothera Gray, 1841 (long-billed partridges)
Perdix Brisson, 1760 (true partridges)
Tribe Phasianini
Syrmaticus Wagler, 1832 (long-tailed pheasants)
Chrysolophus Gray, 1834 (ruffed pheasants)
Phasianus Linnaeus, 1758 (true pheasants)
Catreus Cabanis, 1851 (cheer pheasant)
Crossoptilon Hodgson, 1838 (eared pheasants)
Lophura Fleming, 1822 non Gray, 1827 non Walker, 1856 (gallopheasants)
Phasianinae "Nonerectile clade"
Tribe Pavonini
Rheinardia Maingonnat, 1882 (crested arguses)
Argusianus Rafinesque, 1815 (great argus)
Afropavo Chapin, 1936 (African peafowl)
Pavo Linnaeus, 1758 (Asiatic peafowl)
Polyplectron Temminck, 1807 (peacock-pheasants)
Galloperdix Blyth, 1845 (Indian spurfowls)
Tropicoperdix Blyth, 1859 (chestnut-necklaced and green-legged partridges)
Haematortyx Sharpe, 1879 (crimson-headed partridge)
Tribe Gallini
Bambusicola Gould, 1863 (bamboo partridges)
Gallus Brisson, 1760 (junglefowl, including the domestic chicken)
Peliperdix Bonaparte, 1856 (Latham's francolin)
Ortygornis Reichenbach, 1852 (certain francolins)
Francolinus Stephens, 1819 (certain francolins)
Campocolinus Crowe et al., 2020 (certain francolins)
Scleroptila Blyth, 1852 (certain francolins)
Tribe Coturnicini
Tetraogallus Gray, 1832 (snowcocks)
Ammoperdix Gould, 1851 (sand and see-see partridges)
Synoicus Bosc, 1792 (certain quails)
Margaroperdix Reichenbach, 1853 (Madagascar partridge)
Coturnix Garsault, 1764 (typical Old World quails)
Alectoris Kaup, 1829 (rock partridges)
Perdicula Hodgson, 1837 (bush-quails)
Ophrysia Bonaparte, 1856 (Himalayan quail)
Pternistis Wagler, 1832 (partridge-francolins; African spurfowls)
Past taxonomy
This is the paraphyletic former ordering of Phasianidae, which primarily grouped genera based on appearance and body plans.[12]
Subfamily Perdicinae Horsfield, 1821
Xenoperdix Dinesen et al., 1994 (forest partridges)
Caloperdix Blyth, 1861
Rollulus Bonnaterre, 1791 (crested partridges)
Melanoperdix Jerdon, 1864
Arborophila Hodgson, 1837 (hill partridges)
Rhizothera Gray, 1841
Lerwa Hodgson, 1837
Tropicoperdix Blyth, 1859
Ammoperdix Gould 1851 (see-see and sand partridges)
Synoicus Bosc 1792
Margaroperdix Reichenbach 1853
Coturnix Garsault 1764 (typical Old World quails)
Tetraogallus Gray 1832 (snowcocks)
Alectoris Kaup 1829 (rock partridges)
Pternistis Wagler 1832 (partridge-francolins; African spurfowls)
Ophrysia Bonaparte 1856
Perdicula Hodgson 1837 (bush-quails)
Bambusicola Gould 1863 (bamboo partridges)
Scleroptila Blyth 1852
Peliperdix Bonaparte 1856
Francolinus Stephens 1819 (true francolins)
Ortygornis Reichenbach, 1852
Campocolinus Crowe et al 2020
Perdix Brisson, 1760 (true partridges)
Haematortyx Sharpe, 1879
Galloperdix Blyth, 1845 (Indian spurfowls)
Tetraophasis Elliot, 1871 (monal-partridges)
Subfamily Meleagridinae
Meleagris Linnaeus, 1758 (turkeys)
Subfamily Phasianinae (pheasants, peafowl, junglefowl, monals, and tragopans)
Polyplectron Temminck, 1807 (peacock-pheasants)
Gallus Brisson, 1760 (junglefowl, including the domestic chicken)
Ithaginis Wagler, 1832
Pucrasia Gray, 1841 (koklass pheasant)
Tragopan Cuvier, 1829 non Gray 1841 (tragopans)
Lophophorus Temminck, 1813 non Agassiz, 1846 (monals)
Rheinardia Maingonnat 1882
Argusianus Rafinesque 1815 (argus pheasants)
Afropavo Chapin, 1936 (African peafowl)
Pavo Linnaeus, 1758 (Asiatic peafowl)
Syrmaticus Wagler, 1832 (long-tailed pheasants)
Phasianus Linnaeus, 1758 (true pheasants)
Chrysolophus Gray, 1834 (ruffed pheasants)
Lophura Fleming, 1822 non Gray, 1827 non Walker, 1856 (gallopheasants)
Catreus Cabanis, 1851
Crossoptilon Hodgson, 1838 (eared pheasants)
Subfamily Tetraoninae (grouse)
Bonasa Stephens, 1819 (ruffed grouse)
Tetrastes Keyserling & Blasius, 1840 (hazel grouse)
Centrocercus Swainson 1832 (sage-grouse)
Dendragapus Elliot, 1864 (blue grouse)
Tympanuchus Gloger, 1841 (prairie-chickens and sharp-tailed grouse)
Lagopus Brisson, 1760 (ptarmigans)
Falcipennis Elliot, 1864 (Siberian grouse)
Canachites Stejneger, 1885 (spruce grouse)
Tetrao Linnaeus, 1758 (capercaillies)
Lyrurus Swainson, 1832 (black grouse)
Fossil genera
Extinct genus assignment follows the Mikko's Phylogeny Archive[13] and Paleofile.com websites.[14]
†Alectoris” pliocaena Tugarinov, 1940
†Bantamyx Kuročkin, 1982
†Centuriavis Ksepka, Early, Dzikiewicz & Balanoff, 2022[15]
†Diangallus Hou, 1985
†“Gallus” beremendensis Jánossy, 1976
†“Gallus” europaeus Harrison, 1978
†Lophogallus Zelenkov & Kuročkin, 2010
†Megalocoturnix Sánchez Marco, 2009
†Miophasianus Brodkorb, 1952 [Miophasianus Lambrecht 1933 nomen nudum ; Miogallus Lambrecht 1933]
†Palaeocryptonyx Depéret, 1892 [Chauvireria Boev 1997; Pliogallus Tugarinov 1940b non Gaillard 1939; Lambrechtia Janossy, 1974 ]
†Palaeortyx Milne-Edwards, 1869 [Palaeoperdix Milne-Edwards, 1869]
†Panraogallus Li et al., 2018
†Plioperdix Kretzoi, 1955 [Pliogallus Tugarinov 1940 non Gaillard 1939]
†Rustaviornis Burchak-Abramovich & Meladze, 1972
†Schaubortyx Brodkorb, 1964
†Shandongornis Yeh, 1997
†Shanxiornis Wang et al., 2006
†Tologuica Zelenkov & Kuročkin, 2009
Tribe Tetraonini (grouse)
†Palaealectoris Wetmore, 1930
†Proagriocharis Martin & Tate, 1970
†Rhegminornis Wetmore, 1943
Phylogeny
Cladogram based on a 2021 study by De Chen and collaborators that sequenced DNA flanking ultra-conserved elements. The extinct Himalayan quail (genus Ophrysia) was not included in the study.[16] The species numbers and the inclusion of the genera Canachites, Ortygornis, Campocolinus and Synoicus follows the list maintained by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithologists' Union.[3]
| Phasianidae |
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References
McGowan, P. J. K. (1994). "Family Phasianidae (Pheasants and Partridges)". In del Hoyo, J.; Elliot, A.; Sargatal, J. (eds.). New World Vultures to Guineafowl. Handbook of the Birds of the World. Vol. 2. Barcelona, Spain: Lynx Edicions. pp. 434–479. ISBN 84-87334-15-6.
Harper, D. 1986. Pet Birds for Home and Garden. London: Salamander Books Ltd.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (January 2022). "Pheasants, partridges, francolins". IOC World Bird List Version 12.1. International Ornithologists' Union. Retrieved 6 July 2022.
Johnsgard, P. A. (1986). The Pheasants of the World. Oxford, UK: Oxford University Press.
Johnsgard, P. A. (1988). The Quails, Partridges, and Francolins of the World. Oxford, UK: Oxford University Press.
Kimball, R. T.; Braun, E. L.; Zwartjes, P. W.; Crowe, T. M.; Ligon, J. D. (1999). "A molecular phylogeny of the pheasants and partridges suggests that these lineages are not monophyletic". Molecular Phylogenetics and Evolution. 11 (1): 38–54. doi:10.1006/mpev.1998.0562. PMID 10082609.
Kimball, Rebecca T.; Braun, Edward L. (2014). "Does more sequence data improve estimates of galliform phylogeny? Analyses of a rapid radiation using a complete data matrix". PeerJ. 2: e361. doi:10.7717/peerj.361. PMC 4006227. PMID 24795852.
Mayr, G.; Poshmann, M.; Wuttke, M. (2006). "A nearly complete skeleton of the fossil galliform bird Palaeortyx from the late Oligocene of Germany". Acta Ornithologica. 41 (2): 129–135. doi:10.3161/068.041.0209. S2CID 73586654.
Kimball, R.T.; Hosner, P.A.; Braun, E.L. (2021). "A phylogenomic supermatrix of Galliformes (Landfowl) reveals biased branch lengths". Molecular Phylogenetics and Evolution. 158: 107091. doi:10.1016/j.ympev.2021.107091. ISSN 1055-7903. PMID 33545275. S2CID 231963063.
"Pheasants, partridges, francolins – IOC World Bird List". Retrieved 2022-08-04.
"H&M4 Checklist family by family - The Trust for Avian Systematics". www.aviansystematics.org. Retrieved 2022-08-04.
Çınar, Ümüt (November 2015). "02 → Gᴀʟʟᴏᴀɴsᴇʀᴀᴇ : Gᴀʟʟɪfᴏʀᴍᴇs". English Names of Birds. Archived from the original on 10 August 2016. Retrieved 30 December 2015.
Haaramo, Mikko (2007). "Aves [Avialae]– basal birds". Mikko's Phylogeny Archive. Retrieved 30 December 2015.
"Taxonomic lists- Aves". Paleofile.com (net, info). Retrieved 30 December 2015.
Ksepka, Daniel T.; Early, Catherine M.; Dzikiewicz, Kate; Balanoff, Amy M. (October 2022). "Osteology and neuroanatomy of a phasianid (Aves: Galliformes) from the Miocene of Nebraska". Journal of Paleontology. 97: 223–242. doi:10.1017/jpa.2022.80. ISSN 0022-3360. S2CID 253033983.
Chen, D.; Hosner, P.A.; Dittmann, D.L.; O’Neill, J.P.; Birks, S.M.; Braun, E.L.; Kimball, R.T. (2021). "Divergence time estimation of Galliformes based on the best gene shopping scheme of ultraconserved elements". BMC Ecology and Evolution. 21 (1): 209. doi:10.1186/s12862-021-01935-1. PMC 8609756. PMID 34809586.
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