Pachycephala

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Classis/Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Subclassis/Parvclassis: Neornithes
Infraclassis/Cohors: Neognathae
Cladus: Neoaves
Cladus: Telluraves
Cladus: Australaves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Corvida
Superfamilia: Corvoidea

Familia: Pachycephalidae
Genus: Pachycephala
Species (53): P. albiventris – P. arctitorquis – P. aurea – P. balim – P. caledonica – P. calliope – P. chlorura – P. cinerea – P. citreogaster – P. collaris – P. feminina – P. flavifrons – P. fuliginosa – P. graeffii – P. griseonota – P. homeyeri – P. hyperythra – P. hypoxantha – P. implicata – P. inornata – P. jacquinoti – P. johni – P. lanioides – P. leucogastra – P. lorentzi – P. macrorhyncha – P. melanorhyncha – P. melanura – P. mentalis – P. meyeri – P. modesta – P. monacha – P. nudigula – P. olivacea – P. orioloides – P. ornata – P. orpheus – P. pectoralis – P. phaionota – P. philippinensis – P. richardsi – P. rufiventris – P. rufogularis – P. schlegelii – P. sharpei – P. simplex – P. soror – P. sulfuriventer – P. tenebrosa – P. teysmanni – P. vanikorensis – P. vitiensis – P. utupuae
Name

Pachycephala Vigors, 1825

Typus: Muscicapa pectoralis Latham, 1801 = Pachycephala pectoralis

References
Primary references

Vigors, N.A. 1825. Observations on the Natural Affinities that connect the Orders and Families of Birds. Transactions of the Linnean Society of London 14(3): 395–517. BHL Reference page. p. 444

Additional references

Andersen, M.J., Nyári, A.S., Mason, I.J., Joseph, L., Dumbacher, J.P., Filardi, C.E. & Moyle, R.G. 2014. Molecular systematics of the world’s most polytypic bird: the Pachycephala pectoralis/melanura (Aves: Pachycephalidae) species complex. Zoological Journal of the Linnean Society 170(3): 566–588. DOI: 10.1111/zoj.12088 Paywall Reference page.
Jønsson, K.A., Irestedt, M., Christidis, L., Clegg, S.M., Holt, B.G. & Fjeldså, J. 2014. Evidence of taxon cycles in an Indo-Pacific passerine bird radiation (Aves: Pachycephala). Proceedings of the Royal Society B 281(1777): 20131727. DOI: 10.1098/rspb.2013.1727 ResearchGate Reference page.
Marki, P.Z., Fjeldså, J., Irestedt, M. & Jønsson, K.A. 2018. Molecular phylogenetics and species limits in a cryptically coloured radiation of Australo-Papuan passerine birds (Pachycephalidae: Colluricincla). Molecular Phylogenetics and Evolution 124(5): 100–125. DOI: 10.1016/j.ympev.2018.02.029 Paywall Reference page.

Vernacular names
English: Typical whistlers
español: Silbadores

Pachycephala is a genus of birds native to Oceania and Southeast Asia. They are commonly known as typical whistlers. Older guidebooks may refer to them as thickheads, a literal translation of the generic name, which is derived from the Ancient Greek terms pachys "thick" + kephale "head". This lineage originated in Australo-Papua and later colonized the Indonesian and Philippine archipelagos to the west and the Pacific archipelagos to the east.[1]
Taxonomy

The genus Pachycephala was introduced in 1825 by the Irish zoologist Nicholas Vigors with the Australian golden whistler as the type species.[2][3] The name is derived from the Ancient Greek pakhus meaning "large" or "thick" and kephalē meaning "head".[4]

The genus contains 53 species:[5]

Olive whistler, Pachycephala olivacea
Red-lored whistler, Pachycephala rufogularis
Gilbert's whistler, Pachycephala inornata
Mangrove whistler, Pachycephala cinerea
Green-backed whistler, Pachycephala albiventris
White-vented whistler, Pachycephala homeyeri
Island whistler, Pachycephala phaionota
Biak whistler, Pachycephala melanorhyncha
Rusty whistler, Pachycephala hyperythra
Brown-backed whistler, Pachycephala modesta
Yellow-bellied whistler, Pachycephala philippinensis
Sulphur-vented whistler, Pachycephala sulfuriventer
Bornean whistler, Pachycephala hypoxantha
Vogelkop whistler, Pachycephala meyeri
Grey whistler, Pachycephala simplex
Fawn-breasted whistler, Pachycephala orpheus
Sclater's whistler, Pachycephala soror
Rusty-breasted whistler, Pachycephala calliope (formerly P. fulvotincta)
Selayar whistler, Pachycephala teysmanni (split from P. calliope)
Yellow-throated whistler, Pachycephala macrorhyncha
Baliem whistler, Pachycephala balim
Black-chinned whistler, Pachycephala mentalis
Australian golden whistler, Pachycephala pectoralis
Western whistler, Pachycephala fuliginosa
Bismarck whistler, Pachycephala citreogaster
Oriole whistler, Pachycephala orioloides
Louisiade whistler, Pachycephala collaris
Rennell whistler, Pachycephala feminina
Melanesian whistler, Pachycephala chlorura
New Caledonian whistler, Pachycephala caledonica
White-throated Fiji whistler, Pachycephala vitiensis (formerly Fiji whistler before split)
Yellow-throated Fiji whistler, Pachycephala graeffii (split from P. vitiensis)
Nendo whistler, Pachycephala ornata (split from P. vanikorensis)
Utupua whistler, Pachycephala utupuae (split from P. vanikorensis)
Vanikoro whistler, Pachycephala vanikorensis (formerly Temotu whistler before splits)
Tongan whistler, Pachycephala jacquinoti
Mangrove golden whistler, Pachycephala melanura
Samoan whistler, Pachycephala flavifrons
Hooded whistler, Pachycephala implicata
Bougainville whistler, Pachycephala richardsi
Bare-throated whistler, Pachycephala nudigula
Lorentz's whistler, Pachycephala lorentzi
Regent whistler, Pachycephala schlegelii
Golden-backed whistler, Pachycephala aurea
Rufous whistler, Pachycephala rufiventris
Black-headed whistler, Pachycephala monacha
White-bellied whistler, Pachycephala leucogastra
Babar whistler, Pachycephala sharpei (split from P. macrorhyncha)
Wallacean whistler, Pachycephala arctitorquis
Drab whistler, Pachycephala griseonota
Cinnamon-breasted whistler, Pachycephala johni
White-breasted whistler, Pachycephala lanioides
Morningbird, Pachycephala tenebrosa

Former species

Formerly, some authorities also considered the following species (or subspecies) as species within the genus Pachycephala:

Maroon-backed whistler (as Pachycephala raveni)[6]
Little shrikethrush (fortis) (as Pachycephala fortis)[7]

An unidentified Pachycephala whistler was heard on May 14, 1994 at 1,000 meters ASL south of the summit of Camiguin in the Philippines, where the genus was not previously known to occur. It might have been an undescribed taxon, or simply a vagrant of a known species.[8]

References

Jønsson, K.A. et al. (2014) Evidence of taxon cycles in an Indo-Pacific passerine bird radiation (Aves: Pachycephala). Proc. R. Soc. B.
Vigors, Nicholas Aylward (1825). "Observations on the natural affinities that connect the orders and families of birds". Transactions of the Linnean Society of London. 14 (3): 395–517 [444]. doi:10.1111/j.1095-8339.1823.tb00098.x.
Paynter, Raymond A. Jr, ed. (1986). Check-List of Birds of the World. Vol. 12. Cambridge, Massachusetts: Museum of Comparative Zoology. p. 8.
Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 288. ISBN 978-1-4081-2501-4.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (August 2024). "Whiteheads, sittellas, Ploughbill, Australo-Papuan bellbirds, Shriketit, whistlers". IOC World Bird List Version 14.2. International Ornithologists' Union. Retrieved 20 September 2024.
"Coracornis raveni - Avibase". avibase.bsc-eoc.org. Retrieved 2017-01-29.
"Colluricincla megarhyncha fortis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-02-17.
Balete, Danilo S.; Tabaranza, Blas R. Jr. & Heaney, Lawrence R. (2006) An Annotated Checklist of the Birds of Camiguin Island, Philippines. Fieldiana Zool. New Series 106: 58–72.