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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura

Familia: Megophryidae
Subfamilia: Megophryinae
Genus: Panophrys
Species: Panophrys boettgeri
Name

Panophrys boettgeri (Boulenger, 1899)

Syntypes: BMNH (5 specimens), MCZ 3790.
Type locality: “Kuatun, a village about 270 miles from Foochow, in the mountains at the North-west of the Province of Fokien [= Fujian], at an altitude of 3000 to 4000 feet or more”, China.

Combinations

Leptobrachium boettgeri Boulenger, 1899
Megalophrys boettgeri — Boulenger, 1908
Megophrys boettgeri — Gee & Boring, 1929
Megophrys (Megophrys) boettgeri — Dubois, 1980
Panophrys boettgeri — Rao & Yang, 1997
Megophrys (Xenophrys) boettgeri — Dubois & Ohler, 1998
Xenophrys boettgeri — Ohler, 2003

References
Primary references

Boulenger, 1899, Proc. Zool. Soc. London, 1899: 171.
Ohler, 2003, Alytes, 21: 23.

Links

Frost, D.R. 2021. Amphibian Species of the World: an Online Reference. Version 6.1. Electronic Database accessible at https://amphibiansoftheworld.amnh.org/index.php. American Museum of Natural History, New York, USA. DOI: 10.5531/db.vz.0001 Panophrys boettgeri . Accessed on 11 June 2008.
2007 IUCN Red List of Threatened Species IUCN: Panophrys boettgeri (Least Concern) Downloaded on 11 June 2008.

Vernacular names
English: Boettger's Horned Toad

Boettger's horned toad (Megophrys boettgeri), also known as Boettger's spadefoot toad or the pale-shouldered horned toad, is a species of toad found in southern and southeastern China (the northern border runs roughly from Sichuan in the west to Shanxi in the north and Zhejiang in the east) and north-eastern India (Arunachal Pradesh, Sikkim, and Assam). A closely related but probably as yet undescribed species in found in Tibet.[3] It is not certain that the Indian specimens belong to Megophrys boettgeri either.[1]

The history of this species' discovery is highly international. It was described by George Albert Boulenger, a Belgian zoologist who made his career in the Natural History Museum, London. He named Megophrys boettgeri in honour of Oskar Boettger, a German zoologist, based on specimens collected by Irish ornithologist J. D. La Touche in Guadun village in Wuyishan, Fujian, China.[2]

Male Megophrys boettgeri grow to a snout-vent length of about 36 mm (1.4 in) and females to 43 mm (1.7 in).[4] They are dark grey or brown above, with symmetrical blackish markings and smooth skin with small scattered warts on the head and back.[2]

Megophrys boettgeri is a reasonably common species associated with riparian vegetation, hill streams and leaf-litter in evergreen forest habitats. These frogs breed in streams.[1] Tadpoles are 46 mm (1.8 in) in length.[4]
References

Michael Wai Neng Lau, Geng Baorong, Gu Huiqing, Sushil Dutta, Mohini Mohan Borah, Sabitry Bordoloi (2004). "Megophrys boettgeri". IUCN Red List of Threatened Species. 2004: e.T57632A11667261. doi:10.2305/IUCN.UK.2004.RLTS.T57632A11667261.en. Retrieved 20 November 2021.
Boulenger, George A. (1899). "On a collection of reptiles and batrachians made by Mr. J. D. La Touche in N.W. Fokien, China". Proceedings of the Zoological Society of London. 1899: 159–172.
Frost, Darrel R. (2013). "Xenophrys boettgeri (Boulenger, 1899)". Amphibian Species of the World 5.6, an Online Reference. American Museum of Natural History. Retrieved 21 September 2013.
Fei, L. (1999). Atlas of Amphibians of China (in Chinese). Zhengzhou: Henan Press of Science and Technology. p. 116. ISBN 7-5349-1835-9.

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