Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura
Familia: Microhylidae
Subfamilia: Asterophryinae
Genus: Copiula
Species: C. alpestris – C. annanoreenae – C. bisyllaba – C. derongo – C. exspectata – C. fistulans – C. guttata – C. lennarti – C. major – C. minor – C. mosbyae – C. obsti – C. oxyrhina – C. pipiens – C. rivularis – C. tyleri
Name
Copiula Méhely, 1901
Type species: Phrynixalus oxyrhinus Boulenger, 1898
References
Méhely, 1901, Termés. Füzetek, 24: 193, 242.
Amphibian Species of the World 5.1 Copiula access date 14 June 2008
Vernacular names
English: Mehely Frogs
Copiula is a genus of microhylid frogs endemic to New Guinea. The common name Mehely frogs has been coined for them.[1] They are leaf-litter inhabitants.[2]
Taxonomy
Copiula is probably not monophyletic. Some former Austrochaperina species have already been transferred to this genus, and further ones might follow when more data became available.[1][3]
Species
There are at present 14 species in this genus:[1]
Copiula alpestris (Zweifel, 2000)
Copiula annanoreenae Günther, Richards, and Dahl, 2014
Copiula derongo (Zweifel, 2000)
Copiula exspectata Günther, 2002
Copiula fistulans Menzies and Tyler, 1977 — Lae Mehely Frog
Copiula guttata (Zweifel, 2000)
Copiula lennarti Günther, Richards, and Dahl, 2014
Copiula major Günther, 2002
Copiula minor Menzies and Tyler, 1977 — Milne Bay Mehely Frog
Copiula obsti Günther, 2002
Copiula oxyrhina (Boulenger, 1898) — Misima Island Mehely Frog
Copiula pipiens Burton and Stocks, 1986 — Wirui Mehely Frog
Copiula rivularis (Zweifel, 2000)
Copiula tyleri Burton, 1990 — Sepik Mehely Frog
The AmphibiaWeb[4] reports fewer species, with species that Peloso and colleagues moved in 2016 from Austrochaperina and Oxydactyla missing.[3]
References
Frost, Darrel R. (2016). "Copiula Méhely, 1901". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 20 July 2016.
Zweifel, R. G. (2000). "Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new species". Bulletin of the American Museum of Natural History. 253: 1–130. doi:10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2. hdl:2246/1600.
Peloso, Pedro L.V.; Frost, Darrel R.; Richards, Stephen J.; Rodrigues, Miguel T.; Donnellan, Stephen; Matsui, Masafumi; Raxworthy, Cristopher J.; Biju, S.D.; Lemmon, Emily Moriarty; Lemmon, Alan R.; Wheeler, Ward C. (2016). "The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae)". Cladistics. 32 (2): 113–140. doi:10.1111/cla.12118. S2CID 84925667.
"Microhylidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2016. Retrieved 20 July 2016.
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