Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura
Familia: Microhylidae
Subfamilia: Asterophryinae
Genus: Cophixalus
Species (55): C. aenigma – C. aimbensis – C. albolineatus – C. amabilis – C. ateles – C. australis – C. balbus – C. bewaniensis – C. biroi – C. bombiens – C. cateae – C. caverniphilus – C. cheesmanae – C. clapporum – C. concinnus – C. crepitans – C. cryptotympanum – C. cupricarenus – C. daymani – C. desticans – C. exiguus – C. hannahae – C. hinchinbrookensis – C. hosmeri – C. humicola – C. infacetus – C. interruptus – C. iovaorum – C. kaindiensis – C. kethuk – C. kulakula – C. linnaeus – C. mcdonaldi – C. melanops – C. misimae – C. monosyllabus – C. montanus – C. monticola – C. neglectus – C. nexipus – C. nubicola – C. ornatus – C. pakayakulangun – C. parkeri – C. peninsularis – C. petrophilus – C. phaeobalius – C. pictus – C. pipilans – C. pulchellus – C. rajampatensis – C. riparius – C. salawatiensis – C. saxatilis – C. shellyi – C. sisyphus – C. sphagnicola – C. tagulensis – C. tenuidactylus – C. tetzlaffi – C. timidus – C. tomaiodactylus – C. tridactylus – C. variabilis – C. verecundus – C. verrucosus – C. viridis – C. wempi – C. zweifeli
Name
Cophixalus Boettger, 1892
Type species: Sphenophryne verrucosa Boulenger, 1898
Synonyms
Cophixalus Boettger, 1892
Phrynixalus Boettger, 1895
References
Boettger 1892, Kat. Batr. Samml. Mus. Senckenb. Naturforsch. Ges.: 24.
Günther, R. 2003: First record of the microhylid frog genus Cophixalus from western Papua, Indonesia, with descriptions of two new species (Anura: Microhylidae). Herpetozoa, 16(1-2): 3–21. [not seen]
Günther, R. 2010: Another new Cophixalus species (Amphibia: Anura: Microhylidae) from western New Guinea. Bonn zoological Bulletin, 57(2): 231–240. PDF
Günther, R., Richards, S.J., Tjaturadi, B., Krey, K. 2015. Two new species of the genus Cophixalus from the Raja Ampat Islands west of New Guinea (Amphibia, Anura, Microhylidae). Zoosystematics and Evolution 91(2): 199-213. DOI: 10.3897/zse.91.5411 Open access. Reference page.
Hoskin, C.J. 2004: Australian microhylid frogs (Cophixalus and Austrochaperina): phylogeny, taxonomy, calls, distributions and breeding biology. Australian journal of zoology, 52(3): 237–269. DOI: 10.1071/ZO03056 PDF
Hoskin, C.J. 2013: A new frog species (Microhylidae: Cophixalus) from boulder-pile habitat of Cape Melville, north-east Australia. Zootaxa 3722(1): 61–72. DOI: 10.11646/zootaxa.3722.1.5 Reference page.
Hoskin, C.J.; Aland, K. 2011: Two new frog species (Microhylidae: Cophixalus) from boulder habitats on Cape York Peninsula, north-east Australia. Zootaxa, 3027: 39–51. Preview
Kraus, F. 2012. Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key. Zootaxa 3559: 1–36. PreviewPaywall Reference page.
Kraus, F.; Allison, A. 2009: New species of Cophixalus (Anura: Microhylidae) from Papua New Guinea. Zootaxa, 2128: 1–38. Abstract & excerpt
Kraus, F.; Allison, A. 2009: New microhylid frogs from the Muller Range, Papua New Guinea. ZooKeys, 26: 53–76. DOI: 10.3897/zookeys.26.258
Links
Amphibian Species of the World: an Online Reference. Version 6.0. Cophixalus access date 14 July 2015
ION [needs update]
Nomenclator Zoologicus
Vernacular names
Cophixalus (rainforest frogs or nursery frogs) is a genus of microhylid frogs.[citation needed] These are arboreal species with expanded toe-pads, endemic to Moluccan Islands, New Guinea and northeastern Queensland, Australia.[citation needed]
Species
As of December 2019, Amphibian Species of the World assigns 67 species to the genus Cophixalus.[1]
The source column gives direct links to the sources used:
IUCN description of species at International Union for Conservation of Nature Red List of Threatened Species. IUCN Red List categories are:
EX - Extinct, EW - Extinct in the Wild
CR - Critically Endangered, EN - Endangered, VU - Vulnerable
NT - Near Threatened, LC - Least Concern
DD - Data Deficient, NE - Not Evaluated
ASW description of species at Amphibian Species of the World.
Binomial name and authority | Common name | Geographic range | CS | Source |
---|---|---|---|---|
Cophixalus aenigma Hoskin, 2004 | Thornton rainforest frog | north-eastern Queensland | VU | ASW IUCN |
Cophixalus albolineatus Kraus, 2012 | Mount Shungol, New Guinea | ASW | ||
Cophixalus amabilis Kraus, 2012 | Woodlark Island (PNG) | ASW | ||
Cophixalus ateles (Boulenger, 1898) | Papua rainforest frog | Mount Obree (New Guinea, PNG); possibly widespread in south-eastern Papua New Guinea | DD | ASW IUCN |
Cophixalus australis Hoskin, 2012[2] | southern ornate nursery-frog | north-east Queensland | ASW | |
Cophixalus balbus Günther, 2003 | Yapen rainforest frog | Yapen Island (Indonesia) | DD | ASW IUCN |
Cophixalus bewaniensis Kraus & Allison, 2000 | Menawa rainforest frog | Bewani Mountains (New Guinea, PNG) | DD | ASW IUCN |
Cophixalus biroi (Méhely, 1901) | Sattelberg rainforest frog | northern New Guinea (Indonesia and Papua New Guinea) | LC | ASW IUCN |
Cophixalus bombiens Zweifel, 1985 | buzzing rainforest frog | north-eastern Queensland | NT | ASW IUCN |
Cophixalus cateae Richards and Günther, 2019 | Agogo Range in Southern Highlands Province, Papua New Guinea | ASW | ||
Cophixalus caverniphilus Kraus and Allison, 2009 | Muller Range (New Guinea, PNG) | ASW | ||
Cophixalus cheesmanae Parker, 1934 | Kokoda rainforest frog | eastern Papua New Guinea | LC | ASW IUCN |
Cophixalus clapporum Kraus, 2012 | Woodlark Island (PNG) | ASW | ||
Cophixalus concinnus Tyler, 1979 | rattling rainforest frog | north-eastern Queensland (Australia) | CR | ASW IUCN |
Cophixalus crepitans Zweifel, 1985 | rusty rainforest frog | eastern Cape York Peninsula (Queensland, Australia) | NT | ASW IUCN |
Cophixalus cryptotympanum Zweifel, 1956 | Zweifel's rainforest frog | Mount Dayman (New Guinea, PNG) | DD | ASW IUCN |
Cophixalus cupricarenus Kraus and Allison, 2009 | Rossel Island (PNG) | ASW | ||
Cophixalus daymani Zweifel, 1956 | Dayman rainforest frog | Mount Dayman (PNG) | DD | ASW IUCN |
Cophixalus desticans Kraus and Allison, 2009 | south-easternmost tip of New Guinea, Normanby Island (PNG) | ASW | ||
Cophixalus exiguus Zweifel & Parker, 1969 | Cooktown rainforest frog | north-eastern Queensland (Australia) | NT | ASW IUCN |
Cophixalus hannahae Richards and Günther, 2019 | central cordillera of Papau New Guinea | ASW | ||
Cophixalus hinchinbrookensis Hoskin, 2012[2] | Hinchinbrook Island nursery-frog | Hinchinbrook Island (Queensland, Australia) | ASW | |
Cophixalus hosmeri Zweifel, 1985 | clicking rainforest frog | northern Queensland (Australia) | LC | ASW IUCN |
Cophixalus humicola Günther, 2006 | Waira rainforest frog | Yapen Island, northern New Guinea (Indonesia) | LC | ASW IUCN |
Cophixalus infacetus Zweifel, 1985 | chocolate rainforest frog | north-eastern Queensland (Australia) | LC | ASW IUCN |
Cophixalus interruptus Kraus and Allison, 2009 | Morobe Province (New Guinea, PNG) | ASW | ||
Cophixalus iovaorum Kraus and Allison, 2009 | Mount Obree (New Guinea, PNG) | ASW | ||
Cophixalus kaindiensis Zweifel, 1979 | Kaindi rainforest frog | Mount Kaindi and Mount Missim (New Guinea, PNG) | DD | ASW IUCN |
Cophixalus kethuk Kraus and Allison, 2009 | Rossel Island (PNG) | ASW | ||
Cophixalus kulakula Hoskin and Aland, 2011 | north-eastern Queensland (Australia) | ASW | ||
Cophixalus linnaeus Kraus and Allison, 2009 | Bowutu Mountains (New Guinea, PNG) | ASW | ||
Cophixalus mcdonaldi Zweifel, 1985 | Mcdonald's rainforest frog | north-eastern Queensland (Australia) | EN | ASW IUCN |
Cophixalus melanops Kraus and Allison, 2009 | Tagula Island (PNG) | ASW | ||
Cophixalus misimae Richards & Oliver, 2007 | Misima rainforest frog | Misima Island (PNG) | CR | ASW IUCN |
Cophixalus monosyllabus Günther, 2010 | southern Fakfak Mountains (New Guinea, Indonesia) | ASW | ||
Cophixalus montanus (Boettger, 1895) | mountain rainforest frog | Halmahera Island (Indonesia) | DD | ASW IUCN |
Cophixalus monticola Richards, Dennis, Trenerry, & Werren, 1994 | carbine rainforest frog | northern Queensland (Australia) | EN | ASW IUCN |
Cophixalus neglectus Zweifel, 1962 | brown rainforest frog | northern Queensland (Australia) | EN | ASW IUCN |
Cophixalus nexipus Kraus, 2012 | Mount Obree (New Guinea, PNG) | ASW | ||
Cophixalus nubicola Zweifel, 1962 | Michael rainforest frog | Mount Michael (New Guinea, PNG) | VU | ASW IUCN |
Cophixalus ornatus (Fry, 1912) | ornate rainforest frog | mid-eastern Queensland, Hinchinbrook (Australia) | LC | ASW IUCN |
Cophixalus pakayakulangun Hoskin and Aland, 2011 | north-eastern Queensland (Australia) | ASW | ||
Cophixalus parkeri Loveridge, 1948 | Parker's rainforest frog | central mountainous region of Papua New Guinea | LC | ASW IUCN |
Cophixalus peninsularis Zweifel, 1985 | peninsula rainforest frog | northern Queensland (Australia) | DD | ASW IUCN |
Cophixalus petrophilus Hoskin, 2013 | blotched boulder-frog [3] | |||
Cophixalus phaeobalius Kraus and Allison, 2009 | Bowutu Mountains (New Guinea, PNG) | ASW | ||
Cophixalus pictus Kraus, 2012 | West Papua Province (New Guinea, Indonesia) | ASW | ||
Cophixalus pipilans Zweifel, 1980 | Sempi rainforest frog | from Lae to the Adelbert Mountains (New Guinea, PNG) | LC | ASW IUCN |
Cophixalus pulchellus Kraus & Allison, 2000 | pretty rainforest frog | Mount Hunstein (New Guinea, PNG) | DD | ASW IUCN |
Cophixalus rajampatensis Günther, Richards, Tjaturadi, and Krey, 2015 | Raja Ampat Island group off of western New Guinea (Indonesia) | ASW | ||
Cophixalus riparius Zweifel, 1962 | Wilhelm rainforest frog | central mountain ranges of Papua Guinea (PNG) | LC | ASW IUCN |
Cophixalus salawatiensis Günther, Richards, Tjaturadi, and Krey, 2015 | Raja Ampat Island group off of western New Guinea (Indonesia) | ASW | ||
Cophixalus saxatilis Zweifel & Parker, 1977 | Black Mountain rainforest frog | northern Queensland (Australia) | VU | ASW IUCN |
Cophixalus shellyi Zweifel, 1956 | Shelly's rainforest frog | central mountain ranges of Papua New Guinea, Adelbert Mountains, Huon Peninsula | LC | ASW IUCN |
Cophixalus sphagnicola Zweifel & Allison, 1982 | Morobe rainforest frog | Mount Amungwiwa, Mount Kaini, and Mount Missim (New Guinea, PNG) | LC | ASW IUCN |
Cophixalus tagulensis Zweifel, 1963 | Tagula rainforest frog | Tagula Island (PNG) | DD | ASW IUCN |
Cophixalus tenuidactylus Günther and Richards, 2012 | Kaijende Highlands (New Guinea, PNG) | ASW | ||
Cophixalus tetzlaffi Günther, 2003 | Bomberai rainforest frog | Bomberai Peninsula (New Guinea, Indonesia) | DD | ASW IUCN |
Cophixalus timidus Kraus & Allison, 2006 | Simpson rainforest frog | Mount Simpson (New Guinea, PNG) | DD | ASW IUCN |
Cophixalus tomaiodactylus Kraus and Allison, 2009 | Bowutu Mountains (New Guinea, PNG) | ASW | ||
Cophixalus tridactylus Günther, 2006 | Wondiwoi rainforest frog | Wondiwoi Mountains (New Guinea, Indonesia) | DD | ASW IUCN |
Cophixalus variabilis Kraus & Allison, 2006 | Pekopekowana rainforest frog | south-eastern Papua New Guinea | LC | ASW IUCN |
Cophixalus verecundus Zweifel & Parker, 1989 | Myola rainforest frog | Mount Bellamy (New Guinea, PNG); presumably more widely distributed | DD | ASW IUCN |
Cophixalus verrucosus (Boulenger, 1898) | Moroke rainforest frog | southern and eastern Papua New Guinea | LC | ASW IUCN |
Cophixalus viridis Günther, Richards, and Dahl, 2014 | Green rainforest frog | Muller Range, Western Province, Papua New Guinea | ASW | |
Cophixalus wempi Richards and Oliver, 2010 | Southern Highlands Province (New Guinea, PNG) | ASW | ||
Cophixalus zweifeli Davies & McDonald, 1998 | Cape Melville rainforest frog | northern Queensland (Australia) | DD | ASW IUCN |
References
Frost, Darrel R. 2019. Amphibian Species of the World: an Online Reference. Version 6.0 (10 December 2019). Cophixalus[permanent dead link]. Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.php. American Museum of Natural History, New York, USA. (Accessed: November 17, 2013).
Hoskin, Conrad J. (2012). "Two new frog species (Microhylidae: Cophixalus) from the Australian Wet Tropics region, & redescription of Cophixalus ornatus" (PDF). Zootaxa. 3271: 1–16. doi:10.11646/zootaxa.3271.1.1.
Conrad J. Hoskin (2013). "A new frog species (Microhylidae: Cophixalus) from boulder-pile habitat of Cape Melville, north-east Australia" (PDF excerpt). Zootaxa. 3722 (1): 61–72. doi:10.11646/zootaxa.3722.1.5. PMID 26171515.
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