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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Cladus: Odonatoptera
Cladus: Holodonata
Ordo: Odonata
Subordo: Epiprocta
Infraordo: Anisoptera
Superfamilia: Libelluloidea

Familia: Libellulidae
Subfamilia: Trameinae
Tribus: Rhyothemistini
Genus: Rhyothemis
Species: R. aterrima – R. braganza – R. cognata – R. fenestrina – R. fulgens – R. fuliginosa – R. graphiptera – R. hurleyi – R. mariposa – R. notata – R. obsolescens – R. phyllis – R. plutonia – R. princeps – R. pygmaea – R. regia – R. resplendens – R. rita – R. semihyalina – R. severini – R. splendens – R. triangularis – R. variegata
Name
Rhyothemis Hagen, 1867

Rhyothemis is a genus of dragonfly in the family Libellulidae.[2] They are commonly known as flutterers.

Rhyothemis species are found in Africa, Asia, Australia and the Pacific region.[3] The flight of the genus Rhyothemis is usually fluttering, leading to the common description of "flutterer" for most species in the genus.
Species

The genus Rhyothemis includes the following species:[4]

Male Female Scientific Name Common Name Distribution
Rhyothemis aterrima Selys, 1891 Black Jewel Flutterer Brunei Darussalam; Cambodia; Indonesia (Kalimantan, Sumatera); Malaysia (Sabah, Peninsular Malaysia, Sarawak); Thailand; Viet Nam;
Iridescent Flutterer DSCN1178.jpg
Iridescent Flutterer (5189014461).jpg Rhyothemis braganza Karsch, 1890 Iridescent Flutterer Australia (Western Australia, Queensland, Northern Territory)
Rhyothemis cognata (Rambur, 1842) Madagascar Flutterer Madagascar
Rhyothemis fenestrina (Rambur, 1842) Skylight Flutterer Angola; Benin; Botswana; Cameroon; Congo; Congo, The Democratic Republic of the; Côte d'Ivoire; Gabon; Ghana; Kenya; Liberia; Malawi; Namibia; Nigeria; Rwanda; Senegal; Sierra Leone; Tanzania, United Republic of; Uganda; Zambia
Rhyothemis fulgens Kirby, 1889 Bornean Brown Flutterer Brunei Darussalam; Indonesia (Kalimantan, Sumatera); Malaysia (Peninsular Malaysia, Sarawak); Singapore
Rhyothemis fuliginosa(Male,Japan,2017.09.19,b).jpg
Rhyothemis fuliginosa Selys, 1883 Butterfly Flutterer China, Korea, and Japan
Male Graphic Flutterer up close (15822874480).jpg
Female Graphic Flutterer flank (15387885944).jpg
Rhyothemis graphiptera (Rambur, 1842) Graphic Flutterer[5] or Banded Flutterer or Small Pond Dragonfly Australia (New South Wales, Northern Territory, Queensland, Western Australia); Indonesia (Papua, Lesser Sunda Is., Maluku); New Caledonia; Papua New Guinea
Rhyothemis hurleyi Tillyard, 1926 Indonesia (Papua)
Rhyothemis mariposa Ris, 1913 Mariposa Flutterer[6] Congo, The Democratic Republic of the; Zambia; Angola; Namibia
Rhyothemis notata (Fabricius, 1781) Veiled Flutterer Benin; Cameroon; Central African Republic; Congo; Congo, The Democratic Republic of the; Gabon; Gambia; Ghana; Liberia; Nigeria; Sierra Leone
Rhyothemis obsolescens Kirby, 1889 Bronze Flutterer Brunei Darussalam; China (Hainan); Indonesia (Sumatera, Kalimantan); Malaysia (Sarawak, Sabah, Peninsular Malaysia); Myanmar (Myanmar (mainland)); Philippines; Singapore; Thailand; Viet Nam
Rhyothemis phyllis 2845.jpg
Rhyothemis phyllis (Sulzer, 1776) Yellow-Striped Flutterer Australia (New South Wales, Northern Territory, Queensland); Brunei Darussalam; Cambodia; China; Fiji; Guam; India (Nicobar Is., Andaman Is.); Indonesia (Lesser Sunda Is., Kalimantan, Jawa, Papua, Sumatera, Sulawesi, Maluku); Lao People's Democratic Republic; Malaysia (Sarawak, Sabah, Peninsular Malaysia); Micronesia, Federated States of ; Myanmar (Myanmar (mainland)); New Caledonia; Palau; Papua New Guinea (Papua New Guinea (main island group), North Solomons, Bismarck Archipelago); Philippines; Singapore; Solomon Islands (South Solomons); Thailand; Vanuatu; Viet Nam
Rhyothemis plutonia Selys, 1883 Greater Bluewing or Great Sapphire Flutterer Bangladesh; China (Hainan); India (Assam, West Bengal); Indonesia (Sumatera); Lao People's Democratic Republic; Malaysia (Peninsular Malaysia); Myanmar (Myanmar (mainland)); Nepal; Thailand; Viet Nam
Rhyothemis princeps Kirby, 1894 Australian Sapphire Flutterer or Zircon Flutterer Australia (Queensland); Indonesia (Papua); Papua New Guinea (Papua New Guinea (main island group))
Rhyothemis pygmaea (Brauer, 1867) Small Bronze Flutterer Indonesia (Sulawesi, Maluku, Papua)
Rhyothemis regia (Brauer, 1867) Pied Sapphire Flutterer American Samoa (American Samoa, Swains Is.); Indonesia (Papua, Lesser Sunda Is., Maluku); Malaysia (Sabah); Northern Mariana Islands; Philippines; Samoa; Taiwan, Province of China; Tokelau; Tonga; Wallis and Futuna
Rhyothemis resplendens Selys, 1878 Jewel Flutterer Australia (Queensland); Indonesia (Maluku, Papua); Papua New Guinea (Papua New Guinea (main island group), North Solomons, Bismarck Archipelago); Philippines; Solomon Islands (South Solomons)
Rhyothemis rita Kovacs and Theischinger, 2016 Rita Flutterer Indonesia (Papua)
Rhyothemis semihyalina (Desjardins, 1832) Phantom Flutterer[7] Angola (Cabinda, Angola); Azerbaijan; Benin; Botswana; Cameroon; Chad; Comoros; Congo, The Democratic Republic of the; Côte d'Ivoire; Eritrea; Eswatini; Ethiopia; Gabon; Gambia; Georgia; Ghana; Guinea-Bissau; Iran, Islamic Republic of; Iraq; Kenya; Liberia; Madagascar; Malawi; Mauritius (Mauritius (main island)); Mayotte; Mozambique; Namibia (Namibia (main part), Caprivi Strip); Nigeria; Oman; Rwanda; Réunion; Senegal; Seychelles (Aldabra, Seychelles (main island group)); Sierra Leone; Somalia; South Africa (Free State, Gauteng, KwaZulu-Natal, Limpopo Province, Mpumalanga, North-West Province, Western Cape); South Sudan; Syrian Arab Republic; Tanzania, United Republic of; Togo; Turkey (Turkey-in-Asia); Uganda; Zambia; Zimbabwe
Rhyothemis severini Ris, 1913 Bluetip Flutterer Taiwan, Province of China
Rhyothemis splendens Fraser, 1955 Splendid Flutterer Congo, The Democratic Republic of the
Rhyothemis triangularis Kirby, 1889 Asian Sapphire Flutterer or Lesser Bluewing Brunei Darussalam; China (Fujian, Guangdong); Hong Kong; India (Tamil Nadu, Assam, Kerala, Karnataka); Indonesia (Kalimantan, Jawa, Sumatera); Malaysia (Sarawak, Peninsular Malaysia, Sabah); Philippines; Singapore; Sri Lanka; Taiwan, Province of China

; Thailand

Rhyothemis variegata (Linnaeus, 1763) Common Picture Wing[8] or Variegated Flutterer Bangladesh; Cambodia; China (Guangdong, Hainan, Fujian); Guam; Hong Kong; India (Mahé, Uttaranchal, Daman, Gujarat, Delhi, Haryana, Assam, Dadra-Nagar-Haveli, Pondicherry, Bihar, Jharkand, Goa, Uttar Pradesh, Orissa, Arunachal Pradesh, Yanam, Darjiling, Himachal Pradesh, Sikkim, Andhra Pradesh, Karnataka, Nagaland, Chandigarh, Tripura, Manipur, West Bengal, Maharashtra, Mizoram, Chattisgarh, Meghalaya, Madhya Pradesh, Jammu-Kashmir, Tamil Nadu, Karaikal, Punjab, Andaman Is., Kerala, Diu); Lao People's Democratic Republic; Myanmar (Myanmar (mainland)); Nepal; Sri Lanka; Taiwan, Province of China; Thailand; Viet Nam

References

Hagen, H. (1867). "Die Neuroptera der Insel Cuba". Stettiner Entomologische Zeitung (in German). 28: 215–232 [232] – via Biodiversity Heritage Library.
"Genus Rhyothemis Hagen, 1867". Australian Faunal Directory. Australian Biological Resources Study. 2012. Retrieved 20 March 2017.
Watson, J.A.L.; Theischinger, G.; Abbey, H.M. (1991). The Australian Dragonflies: A Guide to the Identification, Distributions and Habitats of Australian Odonata. Melbourne: CSIRO. p. 278. ISBN 0643051368.
Dennis Paulson; Martin Schorr; Cyrille Deliry. "World Odonata List". University of Puget Sound. Retrieved 15 Feb 2022.
Theischinger, G.; Hawking, J. (2006). The Complete Field Guide to Dragonflies of Australia. CSIRO Publishing. ISBN 0-643-09073-8.
Clausnitzer, V. & Suhling, F. (2010). "Rhyothemis mariposa". IUCN Red List of Threatened Species. 2010: e.T168020A6446785. doi:10.2305/IUCN.UK.2010-3.RLTS.T168020A6446785.en.
Boudot, J.-P.; Clausnitzer, V.; Samraoui, B.; Suhling, F.; Dijkstra, K.-D.B.; Schneider, W. (2016). "Rhyothemis semihyalina". IUCN Red List of Threatened Species. 2016: e.T60037A83869229. doi:10.2305/IUCN.UK.2016-3.RLTS.T60037A83869229.en. Retrieved 12 November 2021.
Subramanian, K. A. (2005). Dragonflies and Damselflies of Peninsular India (PDF).

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