Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Cladus: Odonatoptera
Cladus: Holodonata
Ordo: Odonata
Subordo: Zygoptera
Superfamilia: Lestoidea
Familia: Hemiphlebiidae
Genera: Hemiphlebia – †Electrohemiphlebia – †Jordanhemiphlebia
Name
Hemiphlebiidae Kennedy, 1920
References
Dijkstra, K.-D.B., Kalkman, V.J., Dow, R.A., Stokvis, F.R. & Tol, J. van 2014. Redefining the damselfly families: a comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology 39(1): 68–96. DOI: 10.1111/syen.12035 Open access. Reference page.
Hemiphlebiidae is a family of damselflies, it contains only one extant species, the ancient greenling, native to Southern Australia and Tasmania. The fossil record of the group extends back to the Late Jurassic, making them the oldest known crown group damselflies.[1]
Taxonomy
†Burmahemiphlebia Zheng et al. 2017 Burmese amber, Myanmar, Late Cretaceous (Cenomanian) 99 Ma[2]
†Electrohemiphlebia Lak et al. 2009 Charentese Amber, France, Cretaceous (Albian-Cenomanian) 105-99 Ma[3]
†Enteropia Pritykina and Vassilenko 2014 Shar Teg, Mongolia, Late Jurassic (Tithonian) 145 Ma.[4]
Hemiphlebia Selys, 1869, Australia, Extant.
†Jordanhemiphlebia Kaddumi 2009 Jordanian Amber, Jordan, Early Cretaceous (Albian) 112.6 to 99.7 Ma[3]
†Jurahemiphlebia Bechly, 2019 Solnhofen Limestone, Germany, Late Jurassic (Tithonian) 145 Ma.[1]
†Kachinhemiphlebia Zheng 2020 Burmese amber, Myanmar, Late Cretaceous (Cenomanian) 99 Ma[5]
†Mersituria Vasilenko 2005 Doronino Formation, Russia, Early Cretaceous (Barremian) 130 to 125 Ma.[6]
†Pantelusa Vassilenko 2014 Ora Formation, Israel, Late Cretaceous (Turonian) 94.3 - 89.3 Ma[7]
†Parahemiphlebia Jarzembowski et al. 1998 Durlston Formation, United Kingdom Early Cretaceous (Berriasian) 145-140 Ma[8] Crato Formation, Brazil Early Cretaceous (Aptian) 122-112 Ma.[9]
†Thairia Felker and Vasilenko 2018 Doronino Formation, Russia, Early Cretaceous (Barremian) 130 to 125 Ma.[10]
References
Bechly, Günter (2019-12-20). "New fossil Odonata from the Upper Jurassic of Bavaria with a new fossil calibration point for Zygoptera" (PDF). Palaeoentomology. 2 (6): 618–632–618–632. doi:10.11646/palaeoentomology.2.6.13. ISSN 2624-2834.
Zheng, Daran; Zhang, Qingqing; Nel, André; Jarzembowski, Edmund A.; Zhou, Zhicheng; Chang, Su-Chin; Wang, Bo (2016-05-18). "New damselflies (Odonata: Zygoptera: Hemiphlebiidae, Dysagrionidae) from mid-Cretaceous Burmese amber". Alcheringa: An Australasian Journal of Palaeontology. 41 (1): 12–21. doi:10.1080/03115518.2016.1164402. ISSN 0311-5518. S2CID 89353025.
LAK, MALVINA; FLECK, GÜNTHER; AZAR, DANY; ENGEL FLS, MICHAEL S.; KADDUMI, HANI F.; NERAUDEAU, DIDIER; TAFFOREAU, PAUL; NEL, ANDRÉ (August 2009). "Phase contrast X-ray synchrotron microtomography and the oldest damselflies in amber (Odonata: Zygoptera: Hemiphlebiidae)". Zoological Journal of the Linnean Society. 156 (4): 913–923. doi:10.1111/j.1096-3642.2008.00497.x. ISSN 0024-4082.
L. N. Pritykina and D. V. Vassilenko. 2014. Odonata, in Upper Jurassic Lagerstätte Shar Teg, southwestern Mongolia. Paleontological Journal 48:1641-1647
Zheng, Daran (September 2020). "Odonatans in lowermost Cenomanian Kachin amber: updated review and a new hemiphlebiid damselfly". Cretaceous Research: 104640. doi:10.1016/j.cretres.2020.104640.
D. V. Vasilenko. 2005. New Damselflies (Odonata: Synlestidae, Hemiphlebiidae) from the Mesozoic Transbaikalian Locality of Chernovskie Kopi. Paleontological Journal 39(3):280-283
D. V. Vassilenko. 2014. The first damselfly (Insecta: Odonata, Hemiphlebiidae) recorded from the Turonian of Israel. Far Eastern Entomologist 278:1-7
Jarzembowski, E.A.; Martı́nez-Delclòs, X.; Bechly, G.; Nel, A.; Coram, R.; Escuillié, F. (June 1998). "The Mesozoic non-calopterygoid Zygoptera: description of new genera and species from the Lower Cretaceous of England and Brazil and their phylogenetic significance (Odonata, Zygoptera, Coenagrionoidea, Hemiphlebioidea, Lestoidea)". Cretaceous Research. 19 (3–4): 403–444. doi:10.1006/cres.1997.0113. ISSN 0195-6671.
G. Bechly. 1998. New fossil dragonflies from the Lower Cretaceous Crato Formation of north-east Brazil (Insecta: Odonata). Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) 264:1-66
A. S. Felker and D. V. Vasilenko. 2018. A new genus and species of the damselfly family Hemiphlebiidae from the Lower Cretaceous Chernovskie Kopi locality (eastern Transbaikalia). Paleontological Journal 52:142-145
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