Hellinsia

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Pterophoroidea

Familia: Pterophoridae
Subfamilia: Pterophorinae
Tribus: Oidaematophorini
Genus: Hellinsia
Species:
A

H. acuminatus – H. adumbratus – H. aegyptiacus – H. aethiopicus – H. agassizi – H. agraphodactylus – H. aguilerai – H. albidactylus – H. albilobata – H. aldabrensis – H. alena – H. alfaroi – H. ambo – H. ammonias – H. angela – H. angulofuscus – H. anikini – H. ares – H. argutus – H. arion – H. aruna – H. ascensio – H. auster –
B

H. bahamensis – H. balanotes – H. banosae – H. barbatus – H. basalis – H. basuto – H. batallonica – H. benalcazari – H. beneficus – H. bengtssoni – H. betsiae – H. bhutanensis – H. biangulata – H. bidens – H. bidzilya – H. bifurca – H. bigoti – H. birichevskayae – H. bodneri – H. bogotanus –
C

H. cadmus – H. cajanuma – H. cajas – H. calais – H. caligo – H. callefloridae – H. callidus – H. camposdojordao – H. canari – H. caraca – H. caras – H. carphodactoides – H. carphodactyla – H. carpishia – H. caudelli – H. cervicalis – H. chamelai – H. chewa – H. chionophanes – H. chionoptila – H. chlorias – H. chrysocomae – H. cinerarius – H. citrites – H. cochise – H. colubratus – H. confusus – H. conjunctus – H. conscius – H. contortus – H. coquimboicus – H. corvus – H. cosangae – H. costalba – H. costatus – H. crescens – H. cristobalis – H. cuculla – H. cuenca – H. curvatura – H. curvisacculus –
D

H. delospilus – H. devriesi – H. diana – H. didactylites – H. discors – H. distinctus –
E

H. eborinicolora – H. ecstaticus – H. elena – H. elhacha – H. elliottii – H. emmelinoida – H. emmorus – H. epileucus – H. eros – H. espejoi – H. excors –
F

H. falsus – H. fieldi – H. fishii – H. fissuralba – H. fissuripuncta – H. fletcheri – H. fumiventris – H. furfurosus – H. fusciciliatus – H. fuscomarginata – H. fuscotransversa –
G

H. gielisi – H. glaphyrotes – H. glenni – H. glochinias – H. grandaevus – H. grandis – H. gratiosus – H. griseodactylus – H. gypsotes –
H

H. habecki – H. hamadryadis – H. hami – H. haplistes – H. harpactes – H. haverineni – H. hebrus – H. helianthi – H. hoguei – H. hololeucos – H. homodactylus – H. huayna –
I

H. ignifugax – H. illutus – H. impigritas – H. improbus – H. inconditus – H. innocens – H. inquinatus – H. integratus – H. inulae – H. inulaevorus – H. investis – H. invidiosus – H. iobates – H. iraida – H. iraneaus – H. ishiyamanus –
J

H. jannitae – H. jason – H. joinville – H. jordanica –
K

H. kaiapensis – H. karinupponeni – H. katangae – H. katerina – H. kaweri – H. kellicottii – H. kuwayamai –
L

H. laciniata – H. lacteodactylus – H. lacteolus – H. lenis – H. lienigianus – H. limariae – H. linus – H. llaviuco – H. logistes – H. loja – H. longeanellus – H. longifrons – H. lucayana – H. lumbaquia – H. luteolus –
M

H. madecasseus – H. magnus – H. maldonadoica – H. malesanus – H. mallecoicus – H. mauleicus – H. medius – H. mesoleucus – H. milleri – H. minasgerais – H. mizar – H. mollis – H. mongolicus – H. monserrate – H. monteverda – H. montezerpae – H. montufari – H. morenoi – H. muraseae –
N

H. namizimu – H. nauarches – H. nawrothi – H. nephogenes – H. nigricalcarius – H. nigridactylus – H. nigrosparsus – H. nivalis – H. nodipes – H. novafriburgo – H. novalima – H. nuwara –
O

H. obandoi – H. obscuricilia – H. ochracealis – H. ochricostatus – H. orellanai – H. ossipellis – H. osteodactylus – H. oxyntes –
P

H. paccha – H. pacifica – H. paleaceus – H. pallens – H. palmatus – H. pan – H. papallacta – H. paraglochinias – H. paraguariae – H. paramoi – H. paraochracealis – H. pectodactyla – H. pelodactylus – H. pelospilus – H. perditus – H. phlegmaticus – H. phloeochroa – H. phoebus – H. pichincha – H. pichinguaba – H. pizarroi – H. planiunci – H. podocarpa – H. pollux – H. pondero – H. postnigrata – H. powelli – H. praealtus – H. praenigratus – H. probatus – H. procontias – H. pseudobarbata – H. pseudokorbi – H. punctata – H. puruha – H. purus –
Q

H. quintini – H. quitus –
R

H. rigidus – H. ristoi – H. ruhuruinia – H. ruminahuii –
S

H. salticola – H. sanlucasia – H. saopaulo – H. sapucayae – H. scholasticus – H. scribarius – H. scripta – H. sematias – H. serpens – H. shillongi – H. shyri – H. sichuana – H. simplicissimus – H. siniaevi – H. siskaellus – H. socorroica – H. solanoi – H. sordidatus – H. speideli – H. spermatias – H. sphenites – H. spiculibursa – H. stadias – H. stenoptilioides – H. strauszi – H. sublatus – H. subnotatus – H. subochraceus – H. sucrei – H. sulphureodactylus – H. surinamensis –
T

H. tariensis – H. tenera – H. tephradactyla – H. tepidus – H. tetraonipennis –Hellinsia nawrothi H. thor – H. thoracica – H. timidus – H. tinctus – H. trachyphloeus – H. triadias – H. tripunctatus – H. triton – H. tumeo – H. tupaci –
U

H. ulfi – H. umrani – H. unicolor – H. urbanus –
V

H. varioides – H. varius – H. venapunctus –
W

H. wamenae – H. wojtusiaki – H. wrangeliensis –
X

H. xylodactyla –
Z

H. zakarae – H. zetes
Name

Hellinsia Tutt, 1905

Type species: Pterophorus osteodactylus Zeller, 1841, by original designation.
Synonyms

Leioptilus Wallengren, 1862, homonym, TS: Alucita tephradactyla Hübner, 1813, by subsequent designation by Tutt, 1905. Homonym of Leioptilus Gray, 1860 (Coelenterata).
Utuca Walker, 1864, TS: Utuca ochracealis Walker, 1864, by original designation and monotypy.
Lioptilus Zeller, 1867, emendation and homonym. Homonym of Lioptilus Cabanis, 1850 (Aves).

References

Arenberger, E.; Gurko, V.; Binyason, S.A., 2011: Beitrag zur Kenntnis der Lepidoptera-Fauna des Sudan (Lepidoptera, Pterophoridae). Beiträge zur Entomologie 61(2): 277–284.
Arenberger, E. & J. Wojtusiak, 2001: Pterophoridae aus Venezuela. Quadrifina 4: 65–76. Full article: [1].
Gibeaux, C. 2015a: Note sur les Pterophoridae du Guatémala (Lepidoptera, Pterophoridae, Ochyroticinae, Pterophorinae). Étude des Pterophoridae : 42ème note. Lambillionea, 115(3): 209–216. Reference page.
Gibeaux, C. 2015b. Un Hellinsia Tuft, 1905, nouveau du Guatemala: H. quintini n. sp. (Lepidoptera Pterophoridae Pterophorinae). Antenor 2(1): 106–108. Reference page.
Gielis, C., 2003: World Catalogue of Insects Volume 4, Pterophoroidea & Alucitoidea (Lepidoptera).
Gielis, C., 2003: Review of the Pterophoridae from New Guinea, with descriptions of eight new species (Lepidoptera). Zoologische Mededelingen 77 (21): 349–391. [2]]
Gielis, C., 2003: Review of the Pterophoridae (Lepidoptera) from the Philippines. Zoologische mededelingen 77 (23): 409–424. [3]
Gielis, C., 2008: Ten new species of Afrotropical Pterophoridae (Lepidoptera). Zoologische Mededelingen 82 (1) (2008) [4]
Gielis, C., 2009: On a collection of Pterophoridae (Lepidoptera) from Haut-Katanga, Democratic Republic of the Congo. Revue Suisse de Zoologie 116 (2): 223–256. Abstract [5].
Gielis, C., 2011: Review of the Neotropical species of the family Pterophoridae, Part 2: Pterophorinae (Oidaematophorini, Pterophorini) (Lepidoptera). Zoologische Mededelingen 85 (10): 589–824. Full article: [6].
Gielis, C., 2012: Review of the Neotropical species of the family Pterophoridae, Part 3: Additions from Chile, Ecuador and Paraguay (Lepidoptera). Boletin de la Sociedad Entomológica Aragonesa 51: 105–124.
Gielis, C. 2016. Review of the Neotropical species of the family Pterophoridae, Part 6: additions from Brazil (Lepidoptera). Boletín de la Sociedad Entomológica Aragonesa 58: 33–52. Full article (PDF). Reference page.
Gielis, C. & R. de Vos, 2006, Additions to the Pterophoridae (Lepidoptera) fauna of Papua. Entomofauna 28 (16): 185–200.
Kovtunovich, V.N., 2003: New species of Plum Moths (Lepidoptera: Pterophoridae). Entomological Review 83 (4): 469–473.
Kovtunovich, V.N., Ustjuzhanin, P.Y, de Morney, M., Udovichenko, P. & Bezverkhov, Yu.A. 2017. Fauna of Plume Moths (Lepidoptera: Pterophoridae) of Garden Route National Park, South Africa. Zootaxa 4350(3): 574–582. DOI: 10.11646/zootaxa.4350.3.9. Reference page.
Kovtunovich, V. ; P. Ustjuzhanin ; O.G. Gorbunov & A. Kemal, 2011: On the fauna of the plume moths (Lepidoptera: Pterophoridae) of Ethiopia. Amurian Zoological Journal 3 (4): 354–357. Full article: [7].
Kovtunovich, V. & P. Ustjuzhanin, 2011: On the fauna of the plume moths (Lepidoptera: Pterophoridae) of Lesotho. African Invertebrates 52 (1): 167–175. Abstract: [8].
Kovtunovich, V.N., Ustjuzhanin, P.Y., Pototski, A. & Streltzov, A.N. 2022a. Two new species of the genus Hellinsia Tuff, 1905 (Lepidoptera: Pterophoridae) from South Africa. [[ISSN 1026-051X|Far Eastern Entomologist 455: 1-6. DOI: 10.25221/fee.455.1. Open access. Full article (PDF). Reference page.
Kovtunovich, V.N., Ustjuzhanin,P.Y., Streltzov, A.N. & Ustjuzhanina, A.K. 2022. New data on the Pterophoridae fauna (Lepidoptera) of Uganda with description of a new species. Ecologica Montenegrina 50: 52-55. DOI: 10.37828/em.2022.50.9. Open access. Reference page.
Mano, T. & Y. Yoshiyasu, 2013: A new species of Hellinsia (Lepidoptera, Pterophoridae) from Japan, feeding on Melanthera species (Compositae) with a note on the life history. Lepidoptera Science 64 (1): 36–41.
Matthews, D.L. 2010: A new species of Hellinsia from the southeastern United States (Lepidoptera: Pterophoridae). Bulletin of the Allyn Museum 161: 1–13. Full article (PDF). Reference page.
Ustjuzhanin, P., Kovtunovich, V., Anikin, V. & Aarvik, L. 2016. Additions to the fauna of plume moths (Lepidoptera, Pterophoridae) of Uganda. Zootaxa 4132(3): 393–402. DOI: 10.11646/zootaxa.4132.3.7. Reference page.
Ustjuzhanin, P.Ya. & Kovtunovich, V.N. 2019. New species of plume moths from South Africa (Lepidoptera: Pterophoridae). SHILAP Revista de Lepidopterología 47(186): 379-381. Reference page.
Ustjuzhanin, P., Kovtunovich, V. & Sáfián, S. 2017. New data on the Pterophoridae fauna of Liberia (Lepidoptera: Pterophoridae). Zootaxa 4247(2): 194–200. DOI: 10.11646/zootaxa.4247.2.13. Reference page.
Ustjuzhanin, P.Y., Kovtunovich, V.N., Saldaitis, A. & Streltzov, A.N. 2021a. New species of plume moths (Lepidoptera: Pterophoridae) from China, Sichuan Province. Russian Entomological Journal 30(3): 323–327. DOI: 10.15298/rusentj.30.3.10. Open access. Full article (PDF). Reference page.
Ustjuzhanin, P.Y., Kovtunovich, V.N., Streltzov, A.N. & Ustjuzhanina, A.K. 2022. Two new species of plume moths of the genus Hellinsia Tutt, 1905 (Lepidoptera, Pterophoridae) from Paraguay. Ecologica Montenegrina 54: 12-17. Reference page.
Ustjuzhanin, P.Ya., Kovtunovich, V.N. & Ustjuzhanina, A.K., 2015a: A new species of plume moth (Lepidoptera: Pterophoridae) from Jordan. Zoology in the Middle East 61(1): DOI: 10.1080/09397140.2015.1020619. Reference page.

Hellinsia is a genus of moths in the family Pterophoridae.
Species

The genus contains the following species:

Hellinsia acuminatus (Meyrick, 1920)
Hellinsia adumbratus (Walsingham, 1881)
Hellinsia aegyptiacus (Rebel, 1914)
Hellinsia aethiopicus (Amsel, 1963)
Hellinsia agraphodactylus (Walker, 1864)
Hellinsia aguilerai Gielis, 2011
Hellinsia aistleitneri Arenberger, 2006
Hellinsia albidactylus (Yano, 1963)
Hellinsia albilobata (McDunnough, 1939)
Hellinsia aldabrensis (T.B. Fletcher, 1910)
Hellinsia alfaroi Gielis, 2011
Hellinsia ammonias (Meyrick, 1909)
Hellinsia angela Gielis, 2011
Hellinsia angulofuscus (Gielis, 1991)
Hellinsia ares (Barnes & Lindsey, 1921)
Hellinsia argutus
Hellinsia arion (Barnes & Lindsey, 1921)
Hellinsia aruna Arenberger, 1991
Hellinsia auster (Barnes & Lindsey, 1921)
Hellinsia balanotes (Meyrick, 1908) (sometimes placed in Oidaematophorus)
Hellinsia barbatus
Hellinsia basalis (Möschler, 1890)
Hellinsia basuto Kovtunovich & Ustjuzhanin, 2011[2]
Hellinsia batallonica
Hellinsia bawana Arenberger, 2010
Hellinsia benalcazari Gielis, 2011
Hellinsia beneficus (Yano & Heppner, 1983)
Hellinsia bengtssoni Gielis, 2009
Hellinsia betsiae (Gielis, 1991)
Hellinsia bhutanensis
Hellinsia biangulata Gielis & de Vos, 2006
Hellinsia bigoti (Rougeot, 1983)
Hellinsia bogotanus (Felder & Rogenhofer, 1875)
Hellinsia brandbergi Arenberger, 2004
Hellinsia cadmus (Barnes & Lindsey, 1921)
Hellinsia cajanuma Gielis, 2011
Hellinsia calais (Meyrick, 1930)
Hellinsia callidus (Meyrick, 1913)
Hellinsia canari Gielis, 2011
Hellinsia caras Gielis, 2011
Hellinsia carphodactoides Gielis, 2003
Hellinsia carphodactyla (Hübner, 1813)
Hellinsia caudelli (Dyar, 1903)
Hellinsia cervicalis (Meyrick, 1932)
Hellinsia chamelai (Gielis, 1992)
Hellinsia chlorias (Meyrick, 1908)
Hellinsia chrysocomae (Ragonot, 1875)
Hellinsia chuncheonensis Kim, 2009
Hellinsia cinerarius (Philippi, 1864)
Hellinsia citrites (Meyrick, 1908)
Hellinsia cochise (Barnes & Lindsey, 1921)
Hellinsia colubratus (Meyrick, 1909)
Hellinsia confusus (Braun, 1930)
Hellinsia conjunctus (Zeller, 1877)
Hellinsia conscius (Meyrick, 1920)
Hellinsia contortus (McDunnough, 1938)
Hellinsia conyzae (Gibeaux, 1994)
Hellinsia coquimboicus (Gielis, 1991)
Hellinsia corvus (Barnes & Lindsey, 1921)
Hellinsia costalba
Hellinsia costatus (Barnes & Lindsey, 1921)
Hellinsia crescens (Meyrick, 1926)
Hellinsia cristobalis (B. Landry & Gielis, 1992)
Hellinsia cuculla Gielis, 2011
Hellinsia cyrtoacanthaus Kim, 2009
Hellinsia delospilus (Meyrick, 1921)
Hellinsia devriesi (B. Landry & Gielis, 1992)
Hellinsia didactylites (Ström, 1783)
Hellinsia discors (Meyrick, 1913)
Hellinsia distinctus (Herrich-Schäffer, 1855)
Hellinsia elhacha
Hellinsia elliottii (Fernald, 1893)
Hellinsia emmelinoida Gielis, 2008
Hellinsia emmorus (Walsingham, 1915)
Hellinsia epileucus (Walsingham, 1915)
Hellinsia eros (Barnes & Lindsey, 1921)
Hellinsia excors (Meyrick, 1930)
Hellinsia falsus (Barnes & Lindsey, 1921)
Hellinsia fieldi (Wright, 1921)
Hellinsia fishii (Fernald, 1893)
Hellinsia fissuralba
Hellinsia fissuripuncta
Hellinsia fletcheri
Hellinsia fumiventris (Zeller, 1877)
Hellinsia furfurosus (Meyrick, 1911)
Hellinsia fusciciliatus (Zeller, 1877)
Hellinsia fuscomarginata
Hellinsia fuscotransversa
Hellinsia glaphyrotes (Meyrick, 1908)
Hellinsia glenni (Cashatt, 1972)
Hellinsia glochinias (Meyrick, 1908)
Hellinsia grandaevus (Meyrick, 1931)
Hellinsia grandis (Fish, 1881)
Hellinsia gratiosus (Fish, 1881)
Hellinsia gypsotes (Meyrick, 1937)
Hellinsia habecki Matthews, 2010
Hellinsia haplistes (Meyrick, 1936)
Hellinsia harpactes (Meyrick, 1908)
Hellinsia hebrus (Meyrick, 1932)
Hellinsia helianthi (Walsingham, 1880)
Hellinsia hoguei
Hellinsia hololeucos (Zeller, 1874)
Hellinsia homodactylus (Walker, 1864)
Hellinsia huayna Gielis, 2011
Hellinsia ignifugax (Walsingham, 1915)
Hellinsia illutus (Meyrick, 1917)
Hellinsia improbus
Hellinsia inconditus
Hellinsia innocens
Hellinsia inquinatus
Hellinsia integratus
Hellinsia inulae
Hellinsia inulaevorus (Gibeaux, 1989)
Hellinsia investis
Hellinsia invidiosus
Hellinsia iobates (Barnes & Lindsey, 1921)
Hellinsia iraneaus (Diakonoff, 1952)
Hellinsia ishiyamanus
Hellinsia kaiapensis Gielis, 2003
Hellinsia katangae Gielis, 2009
Hellinsia kellicottii (Fish, 1881)
Hellinsia kuwayamai
Hellinsia laciniata
Hellinsia lacteodactylus
Hellinsia lacteolus
Hellinsia lenis
Hellinsia lienigianus
Hellinsia linus (Barnes & Lindsey, 1921)
Hellinsia logistes
Hellinsia longifrons
Hellinsia lumbaquia Gielis, 2011
Hellinsia luteolus (Barnes & Lindsey, 1921)
Hellinsia madecasseus (Bigot, 1964)
Hellinsia magnus Gielis, 2011
Hellinsia maldonadoica Gielis, 2011
Hellinsia malesanus
Hellinsia mallecoicus
Hellinsia mauleicus
Hellinsia medius (Barnes & Lindsey, 1921)
Hellinsia mesoleucus (Diakonoff, 1952)
Hellinsia milleri Gielis, 2011
Hellinsia mizar (Barnes & Lindsey, 1921)
Hellinsia mollis
Hellinsia mongolicus
Hellinsia monserrate
Hellinsia monteverda
Hellinsia montezerpae
Hellinsia montufari Gielis, 2011
Hellinsia morenoi Gielis, 2011
Hellinsia nauarches
Hellinsia nephogenes
Hellinsia nigricalcarius
Hellinsia nigridactylus
Hellinsia nigrosparsus
Hellinsia nivalis
Hellinsia nodipes
Hellinsia nuwara
Hellinsia obandoi
Hellinsia obscuricilia
Hellinsia ochracealis
Hellinsia ochricostatus
Hellinsia orellanai Gielis, 2011
Hellinsia ossipellis
Hellinsia osteodactyla (Zeller, 1841)
Hellinsia oxyntes
Hellinsia paccha Gielis, 2011
Hellinsia paleaceus
Hellinsia pallens Gielis, 2011
Hellinsia palmatus
Hellinsia pan (Barnes & Lindsey, 1921)
Hellinsia papallacta Gielis, 2011
Hellinsia paraglochinias
Hellinsia paramoi
Hellinsia paraochracealis
Hellinsia pectodactyla
Hellinsia pelospilus
Hellinsia perditus (Barnes & Lindsey, 1921)
Hellinsia phlegmaticus
Hellinsia phloeochroa
Hellinsia phoebus (Barnes & Lindsey, 1921)
Hellinsia pichincha Gielis, 2011
Hellinsia pizarroi Gielis, 2011
Hellinsia pollux (Barnes & Lindsey, 1921)
Hellinsia postnigrata Gielis, 2011
Hellinsia powelli
Hellinsia praealtus
Hellinsia praenigratus
Hellinsia probatus
Hellinsia procontias
Hellinsia pseudobarbata
Hellinsia pseudokorbi
Hellinsia punctata Gielis, 2009
Hellinsia puruha Gielis, 2011
Hellinsia quitus Gielis, 2011
Hellinsia rigidus
Hellinsia ruminahuii Gielis, 2011
Hellinsia scholasticus
Hellinsia scribarius
Hellinsia scripta
Hellinsia sematias
Hellinsia shillongi Kovtunovich, 2003
Hellinsia shyri Gielis, 2011
Hellinsia sichuana
Hellinsia simplicissimus
Hellinsia siniaevi Kovtunovich, 2003
Hellinsia siskaellus
Hellinsia socorroica
Hellinsia solanoi
Hellinsia sordidatus
Hellinsia speideli Gielis, 2003
Hellinsia spermatias
Hellinsia sphenites
Hellinsia spiculibursa
Hellinsia stadias
Hellinsia sublatus
Hellinsia subnotatus
Hellinsia subochraceus
Hellinsia sucrei Gielis, 2011
Hellinsia sulphureodactylus
Hellinsia surinamensis
Hellinsia tariensis Gielis, 2003
Hellinsia tephradactyla (Hübner, 1813)
Hellinsia tepidus
Hellinsia tetraonipennis
Hellinsia thor
Hellinsia thoracica
Hellinsia tinctus
Hellinsia triadias
Hellinsia tripunctatus (=Hellinsia serpens)
Hellinsia triton (Barnes & Lindsey, 1921)
Hellinsia tupaci Gielis, 2011
Hellinsia umrani Kovtunovich, 2003
Hellinsia unicolor
Hellinsia urbanus
Hellinsia varioides
Hellinsia varius (Barnes & Lindsey, 1921)
Hellinsia venapunctus (Barnes & Lindsey, 1921)
Hellinsia wamenae Gielis, 2003
Hellinsia wrangeliensis
Hellinsia zetes

Former species

Hellinsia ecstaticus (Meyrick, 1932)
Hellinsia pacifica
Hellinsia purus

References

On the fauna of the plume moths (Lepidoptera, Pterophoridae) of KwaZulu Natal province (South-African Republic)
Kovtunovich, V. & Ustjuzhanin, P. 2011. On the fauna of the plume moths (Lepidoptera: Pterophoridae) of Lesotho. African Invertebrates 52 (1): 167-175."Archived copy". Archived from the original on 2011-10-04. Retrieved 2011-05-11.