Bromiini

Life-forms

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Cladus: Phytophaga
Superfamilia: Chrysomeloidea

Familia: Chrysomelidae
Subfamilia: Eumolpinae
Tribus: Bromiini

Genera assigned to informal sections (after Seeno & Wilcox, 1982):

Section Bromiites: Anidania – Bromius
Section Leprotites: Aloria – Andosia – Aoria – Aulexis – Brevicolaspis – Crowsonia – Damelia – Demotina – Epistamena – Fidia – Goniopleura – Hemiplatys – Hyperaxis – Irenes – Ischyromus – Leprotoides – Malegia – Neculla – Neofidia – Nephrella – Orthaulexis – Parademotina – Piomera – Pseudaoria – Pseudometaxis – Stasimus – Thootes – Trichotheca – Trichoxantha – Xanthonia – Xanthophorus – †Aoriopsis
Section Myochroites: Acrothinium – Apolepis – Aulacolepis – Cellomius – Chalcosicya – Colaspidea – Colaspina – Damasus – Dictyneis – Eurypelta – ?Glyptoscelis – Heterotrichus – Lepina – Lophea – Mecistes – ?Myochrous – Neocles – Neocloides – Pachnephoptrus – Pachnephorus – Pseudolepis – Rhodopaea – Trichochalcea
Section Nerissites: Casmena – Chiridisia – Dicolectes – Lymidus – Nerissidius – Nerissus – Uhehlia
Section Pseudocolaspites: Badenis – Callipta – Cyno – Diconerissus – Didalsus – Echtrusia – Ennodius – Eurysthenes – Macetes – Macrocoma – Mesocolaspis – Monardiella – Nerissella – Pallena – Pausiris – Phortus – Pseudocolaspis – Pseudostola – Semmiona – Sphaerostola – Tanybria
Section Scelodontites: ?Graphops – Heteraspis – Pagellia – Scelodontina – Scelolanka – Syricta
Section Tomyrites (= Subtribus Ebooina): Eboo
Section Trichochryseites: Caspiana – Dermestops – Eka – Eryxia – Lahejia – Parheminodes – Parnops – Trichochrysea

Genera not assigned to a section: Cryocolaspis – Endroedymolpus – Enneaoria – Erythraella – Pseudoxanthus – †Profidia
Name

Bromiini Baly, 1865 (1863)

Type genus: Bromius Chevrolat in Dejean, 1836
Synonymy

Adoxinae Baly, 1863: 146 [use of younger name Bromiini Baly, 1865 conserved over this name (Art. 40.2)]
Adoxitae: Lefèvre, 1885: 109
Adoxini: Jacoby, 1908: 465
Heteraspinae Baly, 1863: 147
Heteraspites: Chapuis, 1874: 282
Heteraspitae: Lefèvre, 1885: 82
Heteraspini: Jacoby, 1908: 387
Note: Oldest name for tribe; an application to the Commission is needed to conserve usage of the well-established name Bromiini Baly, 1865.
Myochroinae Baly, 1865: 433
Myochroites: Chapuis, 1874: 314
Myochroitae: Lefèvre, 1885: 117
Myochroini: Jacoby, 1908: 459
Bromiinae Baly, 1865: 438 [use of family-group name conserved over Adoxini Baly, 1863 (Art. 40.2)]
Bromiites: Chapuis, 1874: 304
Bromiini: Moseyko & Sprecher-Uebersax, 2010: 619
Tomyrites Chapuis, 1874: 264 [permanently invalid (Art. 39): based on preoccupied type genus]
Tomyritae: Lefèvre, 1885: 64
Tomyrina: Reid, 1993: 61
Scelodontites Chapuis, 1874: 266
Scelodontitae: Lefèvre, 1885: 67
Scelodontini: Jacoby, 1908: 382
Leprotites Chapuis, 1874: 268
Leprotitae: Lefèvre, 1885: 71
Leprotini: Jacoby, 1908: 395
Pseudocolaspites Chapuis, 1874: 287
Pseudocolaspitae: Lefèvre, 1885: 84
Pseudocolaspinae: Jacoby, 1900: 499
Pseudocolaspini: Jacoby, 1908: 466
Goniopleurites Chapuis, 1875: 247 [unavailable name; original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Chapuis (1875)]
Eubrachini Jacoby, 1908: 432
Nerissini Kuntzen, 1912: 42
Nerissites: Seeno & Wilcox, 1982: 50
Odontionopini Clavareau, 1914: 63 [permanently invalid (Art. 39): based on preoccupied type genus]
Trichochryseini Clavareau, 1914: 82
Trichochryseites: Seeno & Wilcox, 1982: 50
Cynoini Clavareau, 1914: 109
Lypesthini Chûjô, 1956: 87
Ebooina Reid, 1993: 61 [replacement name for Tomyrites Chapuis, 1874]
Ebooini: Moseyko, Kirejtshuk & Nel, 2010: 118

References
Primary references

Baly, J.S. 1863. An attempt at a classification of the Eumolpidae. The Journal of Entomology 2: 143–163. BHL. Reference page.
Baly, J.S. 1865. Attempt at a classification of the Eumolpidae. (Cont.). The Journal of Entomology 2: 433–442. BHL
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page. ["Bromiini" conserved over "Adoxini" (Art. 40.2)]
Chapuis, F. 1874. Tome dixième. Famille des phytophages. In: Lacordaire, J.T. & Chapuis, F. (eds) Histoire naturelle des insectes. Genera des coléoptères, ou exposé méthodique et critique de tous les genres proposés jusqu'ici dans cet ordre d'insectes. Librairie Encyclopédique de Roret, Paris. iv + 445 pp. BHL Reference page.
Chapuis, F. 1875. Tome onzième. Famille des phytophages. In: Lacordaire, J.T. & Chapuis, F. (eds) Histoire naturelle des insectes. Genera des coléoptères, ou exposé méthodique et critique de tous les genres proposés jusqu'ici dans cet ordre d'insectes. Librairie Encyclopédique de Roret, Paris. 420 pp. BHL. Reference page.
Chûjô, M. 1956. A Taxonomic Study on the Chrysomelidae (Insecta: Coleoptera) from Formosa. Part VIII. Subfamily Eumolpinae. The Philippine Journal of Science 85(1): 1–180. Full article (PDF).
Clavareau, H. 1914. Pars 59: Chrysomelidae: 11. Eumolpinae. In: Junk, W. & Schenkling, S. (eds.). Coleopterorum Catalogus. Volumen XXIV. Chrysomelidae I. Berlin: W. Junk. 215 pp. BHL. Reference page.
Jacoby, M. 1900. Descriptions of New Species and a New Genus: of South American Eumolpidae, with Remarks on some of the Genera. Transactions of the Entomological Society of London 1900: 453–510. DOI: 10.1111/j.1365-2311.1900.tb02719.x Paywall. BHL.
Jacoby, M. 1908. The Fauna of British India, including Ceylon and Burma. Coleoptera. Chrysomelidae. Vol. 1. London: Francis & Taylor. xx + 534 pp., 2 pls. Internet Archive. BHL Reference page.
Kuntzen, H. 1912. Die Eumolpinensubtribus Nerissini (Coleopt. Chrysomelidae). Archiv für Naturgeschichte. Abteilung A 78(2): 42–57. PDF.
Lefèvre, É. 1885. Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis. Mémoires de la Société Royale des Sciences de Liège (2) 11(16): 1–172. BHL Reference page.
Moseyko, A.G., Kirejtshuk, A.G. & Nel, A. 2010. New genera and new species of leaf beetles (Coleoptera: Polyphaga: Chrysomelidae) from Lowermost Eocene French amber. Annales de la Société entomologique de France (n.s.) 46(1–2): 116–123. DOI: 10.1080/00379271.2010.10697645 Open access. Reference page. [as "Adoxini"]
Moseyko, A.G. & Sprecher-Uebersax, E. 2010. Subfamily Eumolpinae Hope, 1840. Pp. 619–643. In: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, Stenstrup, Denmark. 924 pp. ISBN 978-87-88757-84-2. Google Books. Reference page.
Seeno, T.N. & Wilcox, J.A. 1982. Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography 1: 1–221. Google Books (Snippet view only). Reference page. [as "Adoxini"]
Reid, C.A.M. 1993. Eboo, nom. nov.: redescription of type species (Coleoptera: Chrysomelidae: Eumolpinae). The Coleopterists Bulletin 47(1): 61–67. JSTOR Hybrid open access journal. Reference page.

Additional references

Flowers, R.W. 2012. Chalcosicya maya n. sp, a new Mexican species (Coleoptera: Chrysomelidae: Eumolpinae) and its implications for morphology and biogeography. Insecta Mundi (209): 1–9. PDF. Reference page. [as "Adoxini"]
Medvedev, L.N. & Zoia, S. 2001. A contribution to the knowledge of the oriental Chrysomelidae: two new species of Eumolpinae with description of a new genus (Coleoptera). Bollettino di Zoologia Agraria e di Bachicoltura Ser. II. 33(1): 1–8. PDF. Reference page. [as "Adoxini"]
Moseyko, A.G. 2012. Замечания о роде Aphilenia Weise in Reitter, 1889 (Coleoptera, Chrysomelidae, Eumolpinae) с описанием нового вида из России [Comments on the genus Aphilenia Weise in Reitter, 1889 (Coleoptera, Chrysomelidae, Eumolpinae), with a description of a new species from Russia]. Entomologicheskoe Obozrenie 91(1): 99–105. (in Russian). Abstract. [English transl. in: Entomological Review 92(7): 773–781. DOI: 10.1134/S0013873812070044 Paywall.] Reference page. [as "Adoxini"]
Zoia, S. 2001. Endroedymolpus, a new genus with two new species from the South African Eumolpinae (Coleoptera, Chrysomelidae). Entomologica Basiliensia 23: 311–320. PDF. Reference page. [as "Adoxini"]
Zoia, S. 2012. Eumolpinae (Coleoptera: Chrysomelidae) of Socotra Island. Acta Entomologica Musei Nationalis Pragae 52(s2): 449–501. PDF. Reference page. [as "Adoxini"]
Zoia, S. 2010. New data on African Eumolpinae from the collections of the Naturhistorisches Museum in Basel (Coleoptera, Chrysomelidae). Entomologica Basiliensia et Collectionis Frey 32: 323–341. PDF. Reference page. [as "Adoxini"]

Bromiini (or Adoxini) is a tribe of leaf beetles in the subfamily Eumolpinae. The tribe contains approximately 120 genera, which are found worldwide. They are generally thought to be an artificial group, often with a subcylindrical prothorax without lateral ridges and covered with setae or scales.[6]

Nomenclature

The name "Bromiini" is conserved over the older name "Adoxini" because of Article 40(2) of the ICZN, which states: "If ... a family-group name was replaced before 1961 because of the synonymy of the type genus, the substitute name is to be maintained if it is in prevailing usage. A name maintained by virtue of this Article retains its own author but takes the priority of the replaced name, of which it is deemed to be the senior synonym."[2] Bromiini is cited with its own author and date, followed by the date of the replaced name in parentheses: Bromiini Baly, 1865 (1863).
Taxonomy

Following the leaf beetle classification of Seeno and Wilcox (1982), the genera of Bromiini are divided into eight informal groups or "sections": Bromiites, Leprotites, Myochroites, Nerissites, Pseudocolaspites, Scelodontites, Tomyrites and Trichochryseites.[7] In 1993, the section Tomyrites (interpreted as the subtribe "Tomyrina") was given the replacement name "Ebooina" by C.A.M. Reid, as it was based on a preoccupied genus-group name.[4]

In the Catalog of the leaf beetles of America North of Mexico, published in 2003, Myochroites was placed in synonymy with the section Iphimeites in Eumolpini, while Scelodontites was transferred to Typophorini. Of the genera formerly placed in Myochroites, Glyptoscelis and Myochrous were transferred to Iphimeites in Eumolpini, Colaspidea was transferred to Leprotites, while the placement of the remaining genera was not determined.[8]
Genera

These 68 genera belong to the tribe Bromiini:[9][10][11][12][13][14][15][16][17][18][19]

Acrothinium Marshall, 1865
Andosia Weise, 1896
Anidania Reitter, 1889
Aoria Baly, 1863
†Aoriopsis Moseyko, Kirejtshuk & Nel, 2010[20]
Apolepis Baly, 1863
Aulexis Baly, 1863
Brevicolaspis Laporte, 1833
Bromius Chevrolat in Dejean, 1836
Callipta Lefèvre, 1885
Caspiana Lopatin, 1978
Cellomius Lefèvre, 1888
Chalcosicya Blake, 1930[21]
Colaspidea Laporte, 1833
Colaspina Weise, 1893
Cryocolaspis Flowers, 2004[22]
Damasus Chapuis, 1874
Damelia Clark, 1864
Demotina Baly, 1863
Dermestops Jacobson, 1898
Eboo Reid, 1993[4]
Eka Maulik, 1931[23]
Endroedymolpus Zoia, 2001
Enneaoria Tan, 1981
Erythraella Zoia, 2012[24]
Eryxia Baly, 1865[1]
Fidia Motschulsky, 1861 (= Lypesthes Baly, 1863)[25]
Goniopleura Westwood, 1832
Hemiplatys Baly, 1863
Heterotrichus Chapuis, 1874
Hyperaxis Harold, 1874
Irenes Chapuis, 1874
Lahejia Gahan, 1896
Lepina Baly, 1863
Macetes Chapuis, 1874
Macrocoma Chapuis, 1874
Malegia Lefèvre, 1883
Mecistes Chapuis, 1874
Mesocolaspis Jacoby, 1908
Neocles Chapuis, 1874[5]
Neocloides Jacoby, 1898[26]
Neofidia Strother, 2020 (= Fidia Baly, 1863)[25]
Osnaparis Fairmaire, 1889
Pachnephoptrus Reitter, 1892
Pachnephorus Chevrolat in Dejean, 1836
Parademotina Bryant & Gressitt, 1957
Parheminodes Chen, 1940
Parnops Jacobson, 1894
Phortus Weise, 1899
Piomera Baly, 1863
†Profidia Gressitt, 1963[20]
Pseudaoria Jacoby, 1908
Pseudocolaspis Laporte, 1833[27]
Pseudolepis Medvedev & Zoia, 2001
Pseudometaxis Jacoby, 1900
Pseudoxanthus Zoia, 2010[28]
Rhodopaea Gruev & Tomov, 1968
Semmiona Fairmaire, 1885
Scelodonta Westwood, 1838 (= Heteraspis Chevrolat in Dejean, 1836)
Stasimus Baly, 1863
Tanybria Selman, 1963[29]
Thootes Jacoby, 1890
Trichochalcea Baly, 1878
Trichochrysea Baly, 1861
Trichotheca Baly, 1860
Trichoxantha Medvedev, 1992
Xanthonia Baly, 1863
Xanthophorus Jacoby, 1908

This list is incomplete; you can help by adding missing items. (April 2020)

According to BugGuide and ITIS, the genus Graphops has been transferred to the tribe Typophorini, and Glyptoscelis and Myochrous to the tribe Eumolpini.[12][13]
References
Wikimedia Commons has media related to Bromiini.

Baly, J. S. (1865). "Attempt at a classification of the Eumolpidae. (Cont.)". The Journal of Entomology. 2: 433–442.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; Lawrence, John F.; Lyal, Chris H. C.; Newton, Alfred F.; Reid, Chris A. M.; Schmitt, Michael; Ślipiński, S. Adam; Smith, Andrew B. T. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
Baly, J. S. (1863). "An attempt at a classification of the Eumolpidae". The Journal of Entomology. 2: 143–163.
Reid, C. A. M. (1993). "Eboo, nom. nov.: Redescription of Type Species (Coleoptera: Chrysomelidae: Eumolpinae)". The Coleopterists Bulletin. 47 (1): 61–67. JSTOR 4008912.
Chapuis, F. (1874). "Tome dixième. Famille des phytophages". In Lacordaire, J.T.; Chapuis, F. (eds.). Histoire naturelle des Insectes. Genera des coléoptères. Paris: Librairie Encyclopédique de Roret. pp. i–iv, 1–455.
Jolivet, Pierre; Lawrence, John F.; Verma, Krishna K.; Ślipiński, Adam (2014). "2.7.3 Eumolpinae C. G. Thomson, 1859". In Leschen, R.A.B.; Beutel, R.G. (eds.). Handbook of Zoology. Arthropoda: Insecta: Coleoptera: Volume 3: Morphology and Systematics (Phytophaga). Berlin - Boston: Walter de Gruyter. pp. 217–225. doi:10.1515/9783110274462.189. ISBN 978-3-11-027370-0.
Seeno, T.N.; Wilcox, J.A. (1982). "Leaf beetle genera (Coleoptera, Chrysomelidae)". Entomography. 1: 1–221.
Riley, Edward G.; Clark, Shawn M.; Seeno, Terry N. (2003). Catalog of the leaf beetles of America north of Mexico (Coleoptera: Megalopodidae, Orsodacnidae and Chrysomelidae, excluding Bruchinae). Special Publication No. 1. The Coleopterists' Society. ISBN 978-0-9726087-1-8.
Moseyko, A. G.; Sprecher-Uebersax, E. (2010). "Eumolpinae". In Löbl, I.; Smetana, A. (eds.). Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Stenstrup, Denmark: Apollo Books. pp. 619–643. ISBN 978-87-88757-84-2.
Zoia, S. (2001). "Endroedymolpus, a new genus with two new species from the South African Eumolpinae (Coleoptera, Chrysomelidae)" (PDF). Entomologica Basiliensia. 23: 311–320.
Mohamedsaid, M. S. (2004). Catalogue of the Malaysian Chrysomelidae (Insecta: Coleoptera). Pensoft Series Faunistica. 36. Sofia: Pensoft Publishers. pp. 1–239. ISBN 9546422010. ISSN 1312-0174.
"Adoxini Tribe Information". BugGuide.net. Retrieved 2020-04-23.
"Adoxini Report". Integrated Taxonomic Information System. Retrieved 2020-04-23.
Lawrence, J.F.; Slipinski, A. (2013). Australian Beetles Volume 1: Morphology, Classification and Keys. Csiro Publishing. p. 395. ISBN 978-0-643-09728-5. Retrieved 18 July 2020.
Selman, B. J. (1965). "A revision of the Nodini and a key to the genera of Eumolpidae of Africa (Coleoptera: Eumolpidae)" (PDF). Bulletin of the British Museum (Natural History) Entomology. 16 (3): 141–174. doi:10.5962/bhl.part.21864.
Moseyko, A.G. (2020). "Notes on Asiatic Eumolpinae (Coleoptera, Chrysomelidae)". Entomological Review (published 15 January 2021). 100 (6): 843–862. doi:10.1134/S0013873820060123.
Ordóñez-Reséndiz, María Magdalena; López-Pérez, Sara (2021). "Mexican leaf beetles (Coleoptera: Megalopodidae, Orsodacnidae, and Chrysomelidae): new records and checklist". Revista Mexicana de Biodiversidad. 92: e923873. doi:10.22201/ib.20078706e.2021.92.3873.
Sekerka, L. (16 September 2015). "Eumolpinae". Catálogo Taxonômico da Fauna do Brasil. PNUD. Retrieved 24 October 2021.
Bryant, G. E.; Gressitt, J. L. (1957). "Chrysomelidae of Fiji (Coleoptera)". Pacific Science. 11 (1): 2–91. hdl:10125/8611.
Moseyko, Alexey G.; Kirejtshuk, Alexander G.; Nel, Andre (2010). "New genera and new species of leaf beetles (Coleoptera: Polyphaga: Chrysomelidae) from Lowermost Eocene French amber". Annales de la Société Entomologique de France. Nouvelle Série. 46 (1–2): 116–123. doi:10.1080/00379271.2010.10697645.
Flowers, R. Wills (2012). "Chalcosicya maya n. sp, a new Mexican species (Coleoptera: Chrysomelidae: Eumolpinae) and its implications for morphology and biogeography". Insecta Mundi (209): 1–9.
Flowers, R. Wills (2004). "Cryocolaspis, a New Genus: and Species of Eumolpinae (Chrysomelidae) from Costa Rica". The Coleopterists Bulletin. 58 (1): 97–101. doi:10.1649/607. JSTOR 4009896. S2CID 85218941.
Maulik, S. (1931). "Coleoptera, Chrysomelidæ: Eumolpinæ, Galerucinæ and Halticinæ". Transactions of the Linnean Society of London. 2, Zoology. 19 (2): 241–260. doi:10.1111/j.1096-3642.1931.tb00128.x.
Zoia, S. (2012). "Eumolpinae (Coleoptera: Chrysomelidae) of Socotra Island" (PDF). Acta Entomologica Musei Nationalis Pragae. 52 (supplementum 2): 449–501.
Kumari, S. Amritha; Moseyko, A. G.; Strother, M. S.; Prathapan, K. D. (2020). "Neofidia Strother, a new name for Fidia Baly, 1863 and redescription of Fidia kanaraensis (Jacoby, 1895) with a new host record and notes on natural history (Coleoptera, Chrysomelidae, Eumolpinae)". European Journal of Taxonomy. 654 (654): 1–25. doi:10.5852/ejt.2020.654.
Jacoby, M. (1898). "New species of phytophagous Coleoptera from Australia and the Malayan regions". Annales de la Société Entomologique de Belgique. 42: 350–380.
Laporte, F. L. N. de Caumont (1833). "Mémoire sur les divisions du genre Colaspis". Revue Entomologique. 1: 18–25.
Zoia, S. (2010). "New data on African Eumolpinae from the collections of the Naturhistorisches Museum in Basel (Coleoptera, Chrysomelidae)" (PDF). Entomologica Basiliensia et Collectionis Frey. 32: 323–341.
Selman, B.J. (1963). "A reappraisal of the status of the genus Eubrachis (Eumolpidae, Coleoptera), together with a key to the related genera". Annals and Magazine of Natural History. 13. 6 (70): 637–639. doi:10.1080/00222936308651409.