Geophilidae

Life-forms

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Myriapoda
Classis: Chilopoda
Ordo: Geophilomorpha

Surbordo: Adesmata
Superfamilia: Geophiloidea
Familia: Geophilidae
Subfamiliae: Aphilodontinae – Dignathodontinae – Geophilinae – Linotaeniinae

Overview of genera (155) : Acanthogeophilus – Achilophilus – Agathothus – Agnathodon – Algerophilus – Alloschizotaenia – Aphilodon – Apogeophilus – Arctogeophilus – Arenophilus – Aztekophilus – Barrophilus – Bebekium – Bithyniphilus – Brachygonarea – Caliphilus – Cephalodolichus – Cesachile – Cheiletha – Chilenophilus – Chomatophilus – Clinopodes – Condylona – Damothus – Dekanphilus – Dignathodon – Dinogeophilus – Diphyonyx – Dyodesmophilus – Dysmesus – Ecuadoron – Endogeophilus – Eremerium – Eremorus – Erithophilus – Eurygeophilus – Fagetophilus – Filipponus – Folkmanovius – Galliophilus – Garrina – Geomerinus – Geoperingueyia – Geophilus – Geoporophilus – Gnathomerium – Gnathoribautia – Gosipina – Harpacticellus – Henia – Honuaphilus – Hyphydrophilus – Insigniporus – Italophilus – Ketampa – Khroumiriophilus – Kurdistandus – Leptophilus – Lestophilus – Linotaenia – Macronicophilus – Maoriella – Marsicomerus – Mecistauchenus – Mecophilus – Meinertia – Mesogeophilus – Mesoleptodon – Mixophilus – Monognathodon – Nabocodes – Nannocrix – Navajona – Nearia – Necrophloeophagus – Nesidiphilus – Nesogeophilus – Nicopus – Notadenophilus – Novaralius – Nuevona – Oligna – Onychopodogaster – Orinomerium – Orinomus – Orinophilus – Orpheocricus – Osmanophilus – Pachymerellus – Pachymerinus – Pachymerium – Pachymeroides – Pagotaenia – Pandineum – Pandinium – Paraplanes – Peruphilus – Philacrotherium –Philogeonus – Philosogus – Photophilus – Piestophilus – Plateurytion – Pleurogeophilus – Poabius – Poaphilus – Polycricus – Polyechinogaster – Polygonarea – Poratocricus – Portoricona – Proschizotaenia – Protoplanes – Pseudochaetechelyne – Purcellinus – Pycnona – Queenslandophilus – Ribautia – Schendyloides – Schizonampa – Schizonampus – Schizonium – Schizoribautia – Schizotaenia – Scnipaeus – Scnipoeus – Scolioplanes – Scotophilus – Sepedonophilus – Simoleptus – Simophilus – Snipaeus – Snipoeus – Sogona – Steneurytion – Stenotaenia – Strigamia – Stylolaemus – Sundageophilus – Suturodes – Synerium – Taiyuna – Taschkentia – Telocricus – Thylakiophilus – Timpina – Tomotaenia – Tretechthus – Tuoba – Turkophilus – Tweediphilus – Tylonyx – Watophilus – Zelanion – Zelanoides
Name

Geophilidae
References

Bonato, L., Klarner, B., Widyastuti, R. & Scheu, S. 2016. The first geophilid centipedes from Malesia: a new genus with two new species from Sumatra (Chilopoda, Geophilidae). ZooKeys 605: 53–71. DOI: 10.3897/zookeys.605.9338. Reference page.
Dányi, L. & Tuf, I.H. 2016. Out of Africa: The first introduced African geophilomorph centipede record from a European greenhouse (Chilopoda: Geophilidae). Zootaxa 4067(5): 585–588. DOI: 10.11646/zootaxa.4067.5.6. Reference page.
Dányi, L. & Tuf, I.H. 2017. On the identity of Photophilus Folkmanová, 1928: A new generic synonymy (Chilopoda: Geophilomorpha: Geophilidae). Zootaxa 4347(2): 361–370. DOI: 10.11646/zootaxa.4347.2.10. Reference page.
Pereira, L.A. 2015. A further contribution to the knowledge of two inadequately known species of geophilid centipedes from the Andes of South-Central Chile, currently assigned to the genus Plateurytion Attems, 1909 (Chilopoda: Geophilomorpha). Zootaxa 4027(4): 509–537. DOI: 10.11646/zootaxa.4027.4.3. Preview (PDF) Reference page.
Pereira, L.A. 2016. Plateurytion roigi sp. nov., a new geophilid centipede (Myriapoda: Chilopoda: Geophilomorpha) from Southern Chile. Zootaxa 4162(2): 347–360. DOI: 10.11646/zootaxa.4162.2.9. Reference page.
Tuf, I.H. & Dányi, L. 2015. True identity of Folkmanovius paralellus Dobroruka (Chilopoda: Geophilomorpha: Geophilidae). Zootaxa 4058(3): 444–450. DOI: 10.11646/zootaxa.4058.3.11. Preview (PDF) Reference page.

The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea[3] containing the mostly defunct clades Aphilodontidae,[4] Dignathodontidae,[5] Linotaeniidae,[6] Chilenophilinae,[7] and Macronicophilidae.[8]
Description

Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.[9]

Compared to most other families in the suborder Adesmata, this family features a modest number of leg-bearing segments and limited variation in this number within each species.[10] Two species in this family include centipedes with only 29 pairs of legs: Geophilus persephones (29 pairs in the only specimen, a male), and G. richardi (29 or 31 pairs in males and 33 pairs in females).[11][12] In the order Geophilomorpha, only two species include centipedes with fewer leg pairs, both of them in the family Schendylidae.[11]
Genera

This family contains these genera:[1]

Abatorus
Acanthogeophilus
Achilophilus
Agathothus
Agnathodon
Algerophilus
Alloschizotaenia
Aphilodon
Apogeophilus
Arctogeophilus
Arenophilus
Aztekophilus
Barrophilus
Bebekium
Bithyniphilus
Brachygeophilus
Brachygonarea
Caliphilus
Cephalodolichus
Cheiletha
Chileana
Chilenophilus
Chomatophilus
Clinopodes
Condylona
Damothus
Dekanphilus
Dignathodon
Diphyonyx
Dyodesmophilus
Dysmesus
Ecuadoron
Endogeophilus
Eremerium
Eremorus
Erithophilus
Eurygeophilus
Eurytion
Fagetophilus
Filipponus
Galliophilus
Garrina
Geomerinus
Geoperingueyia
Geophilus
Gnathoribautia
Gosipina
Hapleurytion
Harmostela
Harpacticellus
Henia
Horonia
Hovanyx
Hyphydrophilus
Ketampa
Kurdistanius
Lionyx
Macronicophilus
Maoriella
Mecistauchenus
Mecophilus
Mixophilus
Nabocodes
Nannocrix
Navajona
Nesidiphilus
Nesomerium
Nicopus
Nothogeophilus
Oligna
Orinomerium
Ortognathus
Pachymerellus
Pachymerinus
Pachymerium
Pagotaenia
Pandineum
Peruphilus
Philacroterium
Plestophilus
Plateurytion
Pleurogeophilus
Poaphilus
Polycricus
Polygonarea
Porethus
Portoricellus
Portoricona
Proschizotaenia
Pseudofagetophilus
Purcellinus
Pycnona
Queenslandophilus
Ribautia
Schendyloides
Schizonampa
Schizonium
Schizopleres
Schizotaenia
Sepedonophilus
Serrona
Sogona
Steneurytion
Stenotaenia
Strigamia
Stylolaemus
Sundageophilus
Synerium
Synthophilus
Taiyuna
Taschkentia
Telocricus
Timpina
Tretechthus
Tuoba
Tylonyx
Watophilus
Zantaenia
Zygona
Zygophilus

References

"Geophilidae Leach, 1815". Integrated Taxonomic Information System. Retrieved January 2, 2022.
"Geophilidae".
Bonato, Lucio (2014). "Phylogeny of Geophilomorpha (Chilopoda) inferred from new morphological and molecular evidence". Cladistics. 30 (5): 485–507. doi:10.1111/cla.12060. PMID 34794246. S2CID 86204188. Retrieved 27 October 2021.
Calvanese, Victor C. (2019-11-15). "Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily". Zootaxa. 4698 (1): zootaxa.4698.1.1. doi:10.11646/zootaxa.4698.1.1. PMID 32229999. S2CID 209592191. Retrieved 21 December 2021.
"ITIS - Report: Dignathodontidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
"ITIS - Report: Linotaeniidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
Crabill, Ralph Edwin (1954). "A conspectus of the northeastern North American species of Geophilus (Chilopoda Geophilomorpha Geophilidae)". Proceedings of the Entomological Society of Washington. 56: 172–188. Retrieved 23 October 2021 – via Biodiversity Heritage Library.
"ITIS - Report: Macronicophilidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
Minelli, Alessandro (2011-03-21). Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda. Brill. p. 414. ISBN 978-90-04-18826-6. Retrieved 24 October 2021.
Minelli, Alessandro; Bortoletto, Stefano (1988-04-01). "Myriapod metamerism and arthropod segmentation". Biological Journal of the Linnean Society. 33 (4): 323–343. doi:10.1111/j.1095-8312.1988.tb00448.x. ISSN 0024-4066.
Pereira, Luis Alberto (2013-01-01). "Discovery of a second geophilomorph species (Myriapoda: Chilopoda) having twenty-seven leg-bearing segments, the lowest number recorded up to the present in the centipede order Geophilomorpha". Papéis Avulsos de Zoologia. 53 (13): 163–185. doi:10.1590/S0031-10492013001300001. ISSN 1807-0205.
Bonato, Lucio; Minelli, Alessandro; Drago, Leandro; Pereira, Luis Alberto (2015-09-25). "The phylogenetic position of Dinogeophilus and a new evolutionary framework for the smallest epimorphic centipedes (Chilopoda: Epimorpha)". Contributions to Zoology. 84 (3): 237–253. doi:10.1163/18759866-08403004. ISSN 1875-9866..