Cambarus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Eucarida
Ordo: Decapoda
Subordo: Pleocyemata
Infraordo: Astacidea
Superfamilia: Astacoidea

Familia: Cambaridae
Genus: Cambarus
Subgenera (12): C. (Aviticambarus) – C. (Cambarus) – C. (Depressicambarus) – C. (Erebicambarus) – C. (Exilicambarus) – C. (Glareocola) – C. (Hiaticambarus) – C. (Jugicambarus) – C. (Lacunicambarus) – C. (Puncticambarus) – C. (Tubericambarus) – C. (Veticambarus)
Overview of species

C. acanthura – C. aculabrum – C. acuminatus – C. appalachiensis – C. asperimanus – C. bartonii – C. batchi – C. bouchardi – C. brachydactylus – C. buntingi – C. callainus – C. carinirostris – C. carolinus – C. catagius – C. causeyi – C. chasmodactylus – C. chaugaensis – C. clairitae – C. conasaugaensis – C. coosae – C. coosawattae – C. cracens – C. crinipes – C. cryptodytes – C. cumberlandensis – C. cymatilis – C. deweesae – C. diogenes – C. distans – C. dubius – C. englishi – C. extraneus – C. fasciatus – C. fetzneri – C. franklini – C. friaufi – C. gentryi – C. georgiae – C. girardianus – C. graysoni – C. guenteri – C. halli – C. hamulatus – C. harti – C. hatfieldi – C. hazardi – C. hiwasseensis – C. howardi – C. hubbsi – C. hubrichti – C. jonesi – C. laevis – C. latimanus – C. longirostris – C. longulus – C. ludovicianus – C. manningi – C. miltus – C. monongalensis – C. nerterius – C. nodosus – C. obeyensis – C. ornatus – C. ortmanni – C. obstipus – C. parrishi – C. parvoculus – C. polypilosus – C. pristinus – C. pyronotus – C. reburrus – C. reduncus – C. reflexus – C. robustus – C. rusticiformis – C. sciotensis – C. scotti – C. setosus – C. speciosus – C. sphenoides – C. spicatus – C. striatus – C. strigosus – C. tartarus – C. taylori – C. tenebrosus – C. truncatus – C. unestami – C. veitchorum – C. veteranus – C. zophonastes
Name

Cambarus Erichson, 1846: 97

Type species: Astacus bartonii Fabricius, 1798
Fixation: Subsequent designation of Faxon, 1898: 644

References
Primary references

Erichson, W.F. 1846: Uebersicht der Arten der Gattung Astacus. Archiv für Naturgeschichte (Berlin), 12 (1): 86–103. BHL
Faxon, W. 1898: Observations on the Astacidae in the United States National Museum and in the Museum of Comparative Zoology, with descriptions of new species. Proceedings of the United States National Museum, 2O (1136): 643–694, plates 62-70. BHL
Loughman, Z.J., Welsh, S.A. & Thoma, R.F. 2017. Cambarus (C.) appalachiensis, a new species of crayfish (Decapoda: Cambaridae) from the New River Basin of Virginia and West Virginia, USA. Zootaxa 4243(3): 432–454. DOI: 10.11646/zootaxa.4243.3.2. Reference page.
Loughman, Z.J. & Williams, B.W. 2018. Cambarus polypilosus, a new species of stream-dwelling crayfish (Decapoda: Cambaridae) from the Western Highland Rim of Tennessee, USA. Zootaxa 4403(1): 171–185. DOI: 10.11646/zootaxa.4403.1.10 Reference page.
Schuster, G.A. & Taylor, C.A. 2016. Cambarus (Depressicambarus) clairitae, a new species of crayfish (Decapoda: Cambaridae) from Alabama with a review of the halli Group in the subgenus Depressicambarus. Zootaxa 4193(2): 332–346. DOI: 10.11646/zootaxa.4193.2.8. Reference page.
Thoma, R.F., Loughman, Z.J. & Fetzner, J.W. Jr. 2014. Cambarus (Puncticambarus) callainus, a new species of crayfish (Decapoda: Cambaridae) from the Big Sandy River basin in Kentucky, Virginia, and West Virginia, USA. Zootaxa 3900(4): 541–554. DOI: 10.11646/zootaxa.3900.4.5 Reference page.

Additional references

Bouchard, R.W. & J.W. Bouchard 1995: Two new species and subgenera (Cambarus and Orconectes) of crayfishes (Decapoda: Cambaridae) from the eastern United States. Notulae Naturae 471: 1–21.
Bouchard, R.W., & H.H. Hobbs, Jr. 1976: A New Subgenus and Two New Species of Crayfishes of the Genus Cambarus (Decapoda: Cambaridae) from the Southeastern United States. Smithsonian Contributions to Zoology 224: 1–15. Full article: [1].
Jezerinac, R.F. 1993: A new subgenus and species of crayfish (Decapoda: Cambaridae) of the genus Cambarus, with an amended description of the subgenus Lacunicambarus.Proceedings of the Biological Society of Washington 106: 532–544.
Hobbs, H.H., Jr. 1965: A New Crayfish of the Genus Cambarus from Tennessee with an Emended Definition of the Genus (Decapoda, Astacidae). Proceedings of the Biological Society of Washington 79(15): 109–116
Hobbs, H.H., Jr. 1969: On the Distribution and Phylogeny of the Crayfish Genus Cambarus. In Perry C. Holt, Richard L. Hoffman, and C. Willard Hart, Jr., The Distributional History of the Biota of the Southern Appalachians, Part I: Invertebrates. Virginia Polytechnic Institute, Research Division Monograph, 1: 93–178.
Hobbs, H. H. Jr. 1989: An Illustrated Checklist of the American Crayfishes (Decapoda: Astacidae, Cambaridae, and Parastacidae). Smithsonian Institution Press. 480: 1–236. PDF.
Loughman, Z.J. et al. 2013: Cambarus (P.) theepiensis, a new species of crayfish (Decapoda:Cambaridae) from the coalfields region of Eastern Kentucky and Southwestern West Virginia, USA. Zootaxa 3641(1): 63–73. DOI: 10.11646/zootaxa.3641.1.7 Reference page.
Loughman, Z.J. et al. 2013: Cambarus (C.) hatfieldi, a new species of crayfish (Decapoda: Cambaridae) from the Tug Fork River Basin of Kentucky, Virginia and West Virginia, USA. Zootaxa 3750(3): 223–236. DOI: 10.11646/zootaxa.3750.3.3 Reference page.
Looughman, Z.J., Henkanaththegedara, S.M., Fetzner, J.W. Jr & Thoma, R.F. 2017. A case of Appalachian endemism: Revision of the Cambarus robustus complex (Decapoda: Cambaridae) in the Kentucky and Licking River basins of Kentucky, USA, with the description of three new species. Zootaxa 4269(4): 460–494. DOI: 10.11646/zootaxa.4269.4.4. Reference page.
Loughman, Z.J., Welsh, S.A. & Thoma, R.F. 2019. Cambarus fetzneri sp. nov., a new species of burrowing crayfish (Decapoda: Cambaridae) from the Allegheny Mountains of Virginia and West Virginia, USA. Zootaxa 4651(1): 38–50. DOI: 10.11646/zootaxa.4651.1.2 Paywall Reference page.
Perkins, M.A., Williams, B.W. & Russ, W.T. 2019. Cambarus franklini, a new crayfish (Decapoda: Cambaridae) from the Catawba River Basin in western North Carolina, USA. Zootaxa 4568(3): 520–532. DOI: 10.11646/zootaxa.4568.3.6 Paywall Reference page.
Williams, BW., Delekta, E.M. & Loughman, Z.J. 2019. Redescription and circumscription of the Acuminate Crayfish, Cambarus acuminatus Faxon, 1884 (Decapoda: Cambaridae). Zootaxa 4560(1): 40–50. DOI: 10.11646/zootaxa.4560.1.2 Paywall Reference page.

Cambarus is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 cm (2.0 in) up to approximately 15 cm (5.9 in).
Description

The genus Cambarus is the second largest freshwater crayfish genus inhabiting the Northern Hemisphere, with only sixty fewer species than the genus Procambarus.[2] Though Cambarus are varied across species, the two terminal elements that make up the male form I gonopod form ninety degree angles with the central appendage, allowing for their identification. Unlike the genus Procambarus whose first pleopod tends to have three processes at the tip, Cambarus has only one or two.[2] Cambarus reach 17–26 mm (0.67–1.02 in) carapace lengths in their first year, while average adult carapace length ranges from 55–62 mm (2.2–2.4 in).[3] The name Cambarus comes from an alteration of Latin cammarus, meaning "lobster".[citation needed]

As a genus containing nearly 100 species, Cambarus's coloration is variable.[2] Cambarus bartonii is dark brown, while species like Cambarus pauleyi range from subtle to vibrant blues and reds.[2] Other species are light green or grayish in color.[2]
Biogeography

Most species of Cambarus are restricted to the United States and Canada. They are distributed along the eastern coast, extending from New Brunswick to northern Florida.[2] However, the genus extends as far westward as the Rocky Mountains of Wyoming and Colorado, inhabiting a variety of freshwater environments.[4]
Habitat

Cambarus occupy a range of freshwater environments including streams, rivers, lakes, and burrows. Burrowing species of the genus include Cambarus dubius.[2] Cambarus also include many cave-dwelling species, both stygobites and stygophiles.[3] While salinity and temperature changes minimally affect Cambarus, the genus has shown an intolerance to pollution.[5]
Ecology
Diet

Like other crayfish, Cambarus are foragers. Diets are largely plant-based, though Cambarus also consume small marine organisms like molluscs, larvae, tadpoles and amphibian eggs. Cambarus consume small rodents or birds when available.[3] In their first year, Cambarus typically consume 1-4% of their overall body-weight each day.[3] The genus is central to many freshwater food webs as they help maintain water quality through consumption of algae.[5]
Vulnerability

One of the largest crayfish genera, Cambarus includes a sizable number of vulnerable species.[5] Cave-dwellers like Cambarus jonesi are at risk due to their lack of genetic diversity and low population count. Other species like Cambarus veteranus are at risk due to human practices like logging and mining, which increase sediment amounts in freshwater environments.[2] Increased sediment causes these freshwater environments to be uninhabitable, and Cambarus are forced to relocate as a result.[2]
Growth

Molting occurs among Cambarus approximately 5-10 times during their first year, and 3-5 times during subsequent years. Cambarus remain relatively inactive during periods of molting, as the shedding of chitinous exoskeletons leaves them more vulnerable to predation and injury.[3] Many species of Cambarus continue to grow well into adulthood.[3]
Reproduction

Cambarus typically mate in the early spring. Both Cambarus bartonii and Cambarus robustus only mate once during their three-year life span, with females of both carrying fewer eggs than those of the genus Orconectes.[3]
Classification

The genus Cambarus contains around 100 species,[6] many of which are listed on the IUCN Red List.[7] Species in the genus were formerly divided among 12 subgenera.[8] In a 2017 study,[9] these subgenera were found to lack any phylogenetic validity and were therefore eliminated.

Cambarus aculabrum Hobbs & Brown, 1987 CR – Benton County cave crayfish
Cambarus acuminatus Faxon, 1884 LC - Acuminate Crayfish[10]
Cambarus adustus Thoma, Fetzner, Stocker and Loughman, 2016[11] DD - Dusky Mudbug
Cambarus aldermanorum J. E. Cooper and Price, 2010[12] DD
Cambarus andersoni Jones and Eversole, 2015[13] DD
Cambarus angularis Hobbs & R. W. Bouchard, 1994 NT
Cambarus appalachiensis Loughman, Welsh and Thoma, 2017[14] DD
Cambarus asperimanus Faxon, 1914 LC
Cambarus bartonii (Fabricius, 1798) LC – Appalachian Brook Crayfish
Cambarus batchi Schuster, 1973 LC – bluegrass crayfish
Cambarus bouchardi Hobbs, 1970 VU – Big South Fork crayfish
Cambarus brachydactylus Hobbs, 1953 LC
Cambarus brimleyorum Cooper, 2006 DD
Cambarus buntingi R. W. Bouchard, 1973 LC – Bunting crayfish
Cambarus callainus Thoma, Loughman & Fetzner, 2014 [15] - Big Sandy crayfish
Cambarus carinirostris Hay, 1914 LC – Rock Crayfish
Cambarus carolinus (Erichson, 1846) LC
Cambarus catagius Hobbs & Perkins, 1967 DD – Greensboro Burrowing Crayfish
Cambarus causeyi Reimer, 1966 LC
Cambarus chasmodactylus James, 1966 LC – New River crayfish
Cambarus chaugaensis Prins & Hobbs, 1972 LC – Chauga crayfish
Cambarus clairitae Schuster and Taylor, 2016 [16] DD
Cambarus clivosus Taylor, Soucek & Organ, 2006 VU
Cambarus conasaugaensis Hobbs & Hobbs III, 1962 DD
Cambarus coosae Hobbs, 1981 LC
Cambarus coosawattae Hobbs, 1981 NT – Coosawattae crayfish
Cambarus cracens R. W. Bouchard & Hobbs, 1976 EN
Cambarus crinipes R. W. Bouchard, 1973 LC
Cambarus cryptodytes Hobbs, 1941 LC – Dougherty Plain cave crayfish
Cambarus cumberlandensis Hobbs & R. W. Bouchard, 1973 LC – Cumberland crayfish
Cambarus cymatilis Hobbs, 1970 EN – Conasauga blue burrower
Cambarus davidi J. E. Cooper, 2000 LC – Carolina Ladle Crayfish
Cambarus deweesae R. W. Bouchard & Etnier, 1979 LC – valley flame crayfish
Cambarus distans Rhoades, 1944 LC – boxclaw crawfish
Cambarus diupalma Jones and Eversole, 2015[17] DD
Cambarus doughertyensis Cooper & Skelton, 2003 DD - Dougherty burrowing crayfish
Cambarus dubius Faxon, 1884 LC – upland burrowing crayfish
Cambarus ectopistes Loughman & Williams, 2021[18] DD
Cambarus eeseeohensis Thoma, 2005 VU
Cambarus elkensis Jezerinac & Stocker, 1993 VU – Elk River crayfish
Cambarus englishi Hobbs & Hall, 1972 LC
Cambarus extraneus Hagen, 1870 DD – Chickamauga crayfish
Cambarus fasciatus Hobbs, 1981 DD – Etowah crayfish
Cambarus friaufi Hobbs, 1953 LC – hairy crayfish
Cambarus gentryi Hobbs, 1970 LC
Cambarus georgiae Hobbs, 1981 LC – Little Tennessee crayfish
Cambarus girardianus Faxon, 1884 LC
Cambarus graysoni Faxon, 1914 LC – Two-spot crayfish
Cambarus guenteri Loughman, Henkanaththegedara, Fetzner and Thoma, 2017[19] DD
Cambarus halli Hobbs, 1968 LC
Cambarus hamulatus (Cope, 1881) LC – Prickly cave crayfish
Cambarus harti Hobbs, 1981 EN – Piedmont blue burrower
Cambarus hatfeildi Z. J. Loughman, 2013 DD
Cambarus hazardi Loughman, Henkanaththegedara, Fetzner and Thoma, 2017[20] DD
Cambarus hiwasseensis Hobbs, 1981 LC – Hiwassee crayfish
Cambarus hobbsorum J. E. Cooper, 2001 LC – Rocky River crayfish
Cambarus howardi Hobbs & Hall, 1969 LC – Chattahoochee crayfish
Cambarus hubbsi Creaser, 1931 LC
Cambarus hubrichti Hobbs, 1952 DD – Salem cave crayfish
Cambarus hystricosus Cooper & Cooper, 2003 LC
Cambarus jezerinaci Thoma, 2000 DD
Cambarus johni Cooper, 2006 LC
Cambarus jonesi Hobbs & Barr, 1960 LC – Alabama cave crayfish
Cambarus laconensis Buhay & Crandall, 2009 [21] CR - Lacon Exit cave crayfish
Cambarus latimanus (Le Conte, 1856) LC
Cambarus lenati J. E. Cooper, 2000 NT – Broad River crayfish
Cambarus lentiginosus Jones and Eversole, 2016[22] DD
Cambarus longirostris Faxon, 1885 LC
Cambarus longulus Girard, 1852 LC
Cambarus loughmani Foltz II et al., 2018 - Blue Teays mudbug
Cambarus maculatus Hobbs & Pflieger, 1988 LC – freckled crayfish
Cambarus magerae Thoma and Fetzner, 2015[23] DD
Cambarus manningi Hobbs, 1981 LC
Cambarus monongalensis Ortmann, 1905 LC - Monongahela or blue crayfish
Cambarus nerterius Hobbs, 1964 NT – Greenbrier cave crayfish
Cambarus nodosus R. W. Bouchard & Hobbs, 1976 LC
Cambarus obeyensis Hobbs & Shoup, 1947 CR – Obey crayfish
Cambarus obstipus Hall, 1959 LC
Cambarus ortmanni Williamson, 1907 LC – Ortmann mudbug
Cambarus parrishi Hobbs, 1981 DD – Hiwassee headwater crayfish
Cambarus parvoculus Hobbs & Shoup, 1947 LC – mountain midget crayfish
Cambarus pauleyi Loughman, Thoma, Fetzner and Stocker, 2015[24] DD
Cambarus pecki (Hobbs, 1967) [21] EN - phantom cave crayfish
Cambarus polypilosus Loughman & Williams, 2018[25] DD
Cambarus pristinus Hobbs, 1965 DD – pristine crayfish
Cambarus pyronotus R. W. Bouchard, 1978 DD – fireback crayfish
Cambarus reburrus Prins, 1968 LC – French Broad crayfish
Cambarus reduncus Hobbs, 1956 LC
Cambarus reflexus Hobbs, 1981 LC
Cambarus robustus Girard, 1852 LC – big water crayfish
Cambarus rusticiformis Rhoades, 1944 LC – Depression crayfish
Cambarus sciotensis Rhoades, 1944 LC – Teays River crayfish
Cambarus scotti Hobbs, 1981 LC – Chattooga crayfish
Cambarus setosus Faxon, 1889 NT – bristly cave crayfish
Cambarus smilax Loughman, Simon, and Welch, 2011 NE – Greenbrier crayfish
Cambarus speciosus Hobbs, 1981 NT
Cambarus speleocoopi Buhay & Crandall, 2009 [21] EN - Sweet Home Alabama Crayfish
Cambarus sphenoides Hobbs, 1968 LC
Cambarus spicatus Hobbs, 1956 DD - Broad River spiny crayfish
Cambarus stockeri Thoma, 2011[26] DD
Cambarus striatus Hay, 1902 LC – Hay Crayfish
Cambarus strigosus Hobbs, 1981 DD – lean crayfish
Cambarus subterraneus Hobbs III, 1993 CR – Delaware County cave crayfish
Cambarus tartarus Hobbs & M. R. Cooper, 1972 CR – Oklahoma cave crayfish
Cambarus taylori Loughman, Henkanaththegedara, Fetzner and Thoma, 2017[27] DD
Cambarus tenebrosus Hay, 1902 LC – cavespring crayfish
Cambarus theepiensis Loughman, Foltz, Garrison and Welsh, 2013[28] DD
Cambarus truncatus Hobbs, 1981 NT – Oconee burrowing crayfish
Cambarus tuckasegee Cooper & Schofield, 2002 NT
Cambarus unestami Hobbs & Hall, 1969 LC – Blackbarred crayfish
Cambarus veitchorum J. E. Cooper & M. R. Cooper, 1997 CR – White Spring cave crayfish
Cambarus veteranus Faxon, 1914 DD – Guyandotte River crayfish
Cambarus williami R. W. Bouchard & J. W. Bouchard, 1995 NT – Brawleys Fork crayfish
Cambarus zophonastes Hobbs & Bedinger, 1964 CR – Hell Creek cave crayfish

References

Horton H. Hobbs Jr. (1974). "A Checklist of the North and Middle American Crayfishes (Decapoda: Astacidae and Cambaridae)". Smithsonian Contributions to Zoology. 166 (166): 1–161. doi:10.5479/si.00810282.166.
Longshaw, Matt (2016). Biology and Ecology of Crayfish. New York: CRC Press. pp. 17–18. ISBN 9781498767323.
Holdich, David M. (2002). Biology of Freshwater Crayfish. London: Blackwell Science. pp. 609–613. ISBN 978-0-632-05431-2.
Cordeiro, J. (2014). "Cambarus diogenes". Encyclopedia of Life. Retrieved March 29, 2018.
Buhay, J; et al. (February 2007). "Molecular taxonomy in the dark: Evolutionary history, phylogeography, and diversity of cave crayfish in the subgenus Aviticambarus, genus Cambarus". Molecular Phylogenetics and Evolution. 42 (2): 435–448. doi:10.1016/j.ympev.2006.07.014. PMID 16971141.
"Cambarus". Integrated Taxonomic Information System.
"Search". IUCN Red List. IUCN. Retrieved October 7, 2010.
James W. Fetzner Jr. (January 14, 2008). "Genus Cambarus Erichson, 1846". Crayfish Taxon Browser. Carnegie Museum of Natural History. Archived from the original on April 29, 2014. Retrieved March 9, 2010.
Crandall, Keith A; De Grave, Sammy (September 2017). "An updated classification of the freshwater crayfishes (Decapoda: Astacidea) of the world, with a complete species list". Journal of Crustacean Biology. 37 (5): 615–653. doi:10.1093/jcbiol/rux070. ISSN 0278-0372.
Williams, BW; Delekta, EM; Loughman, ZJ (22 February 2019). "Redescription and circumscription of the Acuminate Crayfish, Cambarus acuminatus Faxon, 1884 (Decapoda: Cambaridae)". Zootaxa. 4560 (1): 40–50. doi:10.11646/zootaxa.4560.1.2. PMID 30790990. S2CID 73459024. Retrieved 26 March 2022.
"Cambarus adustus". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus aldermanorum". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus andersoni". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus appalachiensis". Integrated Taxonomic Information System. Retrieved 25 March 2022.
Roger F. Thoma; Zachary J. Loughman; James W. Fetzner Jr. (2014). "Cambarus (Puncticambarus) callainus, a new species of crayfish (Decapoda: Cambaridae) from the Big Sandy River basin in Kentucky, Virginia, and West Virginia, USA" (PDF). Zootaxa. 3900 (4): 541–554. doi:10.11646/zootaxa.3900.4.5. PMID 25543755.
"Cambarus clairitae". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus diupalma". Integrated Taxonomic Information System. Retrieved 25 March 2022.
Loughman, Zachary J; Williams, Bronwyn W (December 2021). "Cambarus ectopistes sp. nov., a new stream-dwelling crayfish (Decapoda: Cambaridae) from the French Broad, Pigeon, and Nolichucky River watersheds in the Appalachian Mountain region of North Carolina and Tennessee, USA". Zootaxa. 5082 (4): 322–340. doi:10.11646/zootaxa.5082.4.2. PMID 35390961. S2CID 245339895. Retrieved 27 March 2022.
"Cambarus guenteri". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus hazardi". Integrated Taxonomic Information System. Retrieved 25 March 2022.
Jennifer E. Buhay; Keith A. Crandall (2009). "Taxonomic revision of cave crayfish in the genus Cambarus subgenus Aviticambraus (Decapoda: Cambaridae) with descriptions of two new species, C. speleocoopi and C. laconensis, endemic to Alabama, U.S.A." Journal of Crustacean Biology. 29 (1): 121–134. doi:10.1651/08-3089.1.
"Cambarus lentiginosus". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus magerae". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus pauleyi". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus polypilosus". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus stockeri". Integrated Taxonomic Information System. Retrieved 25 March 2022.
"Cambarus taylori". Integrated Taxonomic Information System. Retrieved 25 March 2022.

"Cambarus theepiensis". Integrated Taxonomic Information System. Retrieved 25 March 2022.