Trachycarpeae

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Monocots
Cladus: Commelinids
Ordo: Arecales

Familia: Arecaceae
Subfamilia: Coryphoideae
Tribus: Trachycarpeae
Subtribus: Livistoninae – Rhapidinae – Unplaced Trachycarpeae
Name

Trachycarpeae Satake, Hikobia 3: 121. (1962)

Type genus: Trachycarpus H.Wendl., Bull. Soc. Bot. France 8: 429. (1862)

Synonyms

Heterotypic
Livistoneae J.Dransf., N.W.Uhl, Asmussen, W.J.Baker, M.M.Harley & C.Lewis, Kew Bull. 60(4): 561. (2005)
Type genus: Livistona R.Br., Prodr. Fl. Nov. Holl.: 267 (1810)

Note: Unplaced Trachycarpeae a useful category (Baker & Dransfield, 2016) and so is retained here.
References

Satake, Y. 1962. Hikobia; Journal of the Hiroshima Botanical Club. Hiroshima 3: 121.
Baker, W.J. & Dransfield, J. 2016. Beyond Genera Palmarum: progress and prospects in palm systematics. Botanical Journal of the Linnean Society 182(2): 207-233. DOI: 10.1111/boj.12401 Reference page.
Dransfield, J., Uhl, N.W., Asmussen, C.B., Baker, W.J., Harley, M.M. & Lewis, C.E. 2005. A New Phylogenetic Classification of the Palm Family, Arecaceae. Kew Bulletin 60 (4): 559–569. JSTOR Stable PDF. Reference page.
Dransfield, J., Uhl, N.W., Asmussen, C.B., Baker, W.J., Harley, M.M. & Lewis, C.E. 2008. Genera palmarum: The Evolution and Classification of Palms. Kew Publishing: Royal Botanic Gardens, Kew. ISBN 9781842461822 Reference page.

Trachycarpeae is a tribe of palms in subfamily Coryphoideae of the plant family Arecaceae.[1][2] It has the widest distribution of any tribe in Coryphoideae and is found on all continents (except Antarctica), though the greatest concentration of species is in Southeast Asia.[3] Trachycarpeae includes palms from both tropical and subtropical zones; the northernmost naturally-occurring palm is a member of this tribe (Chamaerops humilis).[4] Several genera can be found in cultivation in temperate areas, for example species of Trachycarpus, Chamaerops, Rhapidophyllum and Washingtonia.[5]
Description

Palms in this tribe have palmate leaves with induplicate folds (reduplicate in Guihaia).[3] Plants may be tall, single-stemmed trees (e.g. Copernicia, Brahea, Pritchardia), acaulescent with short, squat trunks (e.g. Maxburretia, Johannesteijsmannia), multi-stemmed (e.g. Rhapis, Acoelorrhaphe) or branched and prostrate (e.g. Serenoa). These palms flower regularly throughout their lives (pleonanthic) and may be dioecious, monoecious or hermaphroditic.[3]
Taxonomy

Trachycarpeae is one of eight tribes in the subfamily Coryphoideae.[6] The tribe is monophyletic, but phylogenetic studies have yet to reveal its closest relatives, though they could be the Phoeniceae, or the Sabaleae and Cryosophileae.[3] Initially described as tribe 'Livistoneae', the name Trachycarpeae has priority.[3] In previous classifications, all the members of this tribe were included in tribe Corypheae.[7]

Trachycarpeae is divided into two subtribes: Rhapidinae have flowers with three separate carpels, whereas in subtribe Livistoniinae the flower carpels are free at the base, but the styles are fused together.[3] All genera in Rhapidinae are native to the Old World, except North American Rhapidophyllum. Livistoninae are widely distributed in both the New World and Southeast Asia and Australia. A single species (Livistona carinensis) occurs in Africa. Several genera in this tribe have yet to be allocated to a subtribe, due to a lack of convincing data from phylogenetic studies.[6]

Rhapidinae[3][6]

Chamaerops - (1 sp., C. humilis, Mediterranean)
Guihaia - (3 spp., China, Vietnam)
Trachycarpus - (11 spp., Himalayas, Indochina)
Rhapidophyllum - (1 sp., R. hystrix, Southeast USA)
Maxburretia - (3 spp., Malay Peninsula)
Rhapis - (11 spp., Indochina, Sumatra)

Livistoninae[3][6]

Livistona - (approx. 27 spp., Southeast Asia, Australia, NE Africa)
Licuala - (167 spp., Southeast Asia)
Lanonia - (13 spp., Indochina)
Johannesteijsmannia - (4 spp., Malaysia, Indonesia)
Pholidocarpus - (6 spp., Malaysia, Indonesia)
Saribus - (9 spp., Philippines to New Caledonia)

Unplaced genera[3][6]

Acoelorrhaphe - (1 sp., A. wrightii, Caribbean)
Serenoa - (1 sp., S. repens, Southeast USA)
Brahea - (approx. 10 spp., Mexico & Guatemala)
Colpothrinax - (3 spp., Central America & Cuba)
Copernicia - (21 spp., Caribbean, South America)
Pritchardia - (approx. 27 spp., Pacific)
Washingtonia - (2 spp., California, Mexico)

The above classification was published prior to the recognition of the genera Saribus and Lanonia; however, both are clearly members of subtribe Livistoniinae. Saribus includes species formerly in Livistona and monotypic Pritchardiopsis,[8] while Lanonia species were previously included in Licuala.[9][10]

References

Baker, William J.; Dransfield, John (2016). "Beyond Genera Palmarum: progress and prospects in palm systematics". Botanical Journal of the Linnean Society. 182 (2): 207–233. doi:10.1111/boj.12401.
Dransfield, John; Uhl, Natalie W.; Asmussen, Conny B.; Baker, William J.; Harley, Madeline M.; Lewis, Carl E. (2008). Genera Palmarum: The Evolution and Classification of Palms. Royal Botanic Gardens, Kew. ISBN 978-1-84246-182-2.
Dransfield, John; Uhl, Natalie W.; Asmussen, Conny B.; Baker, William J.; Harley, Madeline M.; Lewis, Carl E. (2008). Genera Palmarum - The Evolution and Classification of Palms. Royal Botanic Gardens, Kew. ISBN 9781842461822.
"Chamaerops humilis - Plant Finder". www.missouribotanicalgarden.org. Retrieved 2018-03-02.
"Cold Hardy Palms for Temperate Gardens". Plant Delights Nursery. Retrieved 2018-03-02.
Dransfield, John; Uhl, Natalie W.; Asmussen, Conny B.; Baker, William J.; Harley, Madeline M.; Lewis, Carl E. (2005). "A new phylogenetic classification of the palm family, Arecaceae". Kew Bulletin. 60: 559–569 – via ResearchGate.
Uhl, Natalie W.; Dransfield, John (1987). Genera Palmarum: A Classification of Palms Based on the Work of Harold E. Moore, Jr. L.H. Bailey Hortorium. ISBN 9780935868302.
Bacon, Christine D.; Baker, William J. (2011). "Saribus Resurrected". Palms. 55 (3): 109–116 – via ResearchGate.
Henderson, Andrew J.; Bacon, Christine D. (2011). "Lanonia (Arecaceae: Palmae), a New Genus from Asia, with a Revision of the Species". Systematic Botany. 36 (4): 883–895. doi:10.1600/036364411x604903. JSTOR 41416905. S2CID 84318474.
Henderson, Andrew; Dung, Nguyen Quoc (2017-09-29). "New species of Lanonia, Licuala, and Pinanga (Arecaceae) from Vietnam". Phytotaxa. 323 (2): 159. doi:10.11646/phytotaxa.323.2.4. ISSN 1179-3163.