Cydonia oblonga

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordo: Rosales

Familia: Rosaceae
Subfamilia: Amygdaloideae
Tribus: Maleae
Subtribus: Malinae
Genus: Cydonia
Species: Cydonia oblonga
Name

Cydonia oblonga Mill., Gard. Dict. ed. 8: n.º 1 (1768).

References

Miller, P. 1768. Gard. dict. ed. 8: Cydonia no. 1.

Vernacular names
Afrikaans: Kweper
беларуская: Айва
bosanski: Dunja
čeština: kdouloň obecná
Deutsch: Quitte
Ελληνικά: Κυδωνία
English: Quince
español: Membrillo
suomi: Kvitteni
Nordfriisk: Kwitenbuum
français: Coing
Gàidhlig: cuinnse
magyar: birs,birsalma, birskörte
italiano: Cotogno
latviešu: Parastā cidonija
Mirandés: Burmelheiro
Nederlands: Kweepeer
polski: pigwa pospolita
português: Marmeleiro
русский: Айва обыкновенная, Айва продолговатая
slovenčina: dula podlhovastá
svenska: Kvitten
Türkçe: Ayva
українська: Айва
vèneto: Cognaro, Cogno

The quince (/ˈkwɪns/; Cydonia oblonga) is the sole member of the genus Cydonia in the family Rosaceae (which also contains apples and pears, among other fruits). It is a deciduous tree that bears hard, aromatic bright golden-yellow pome fruit, similar in appearance to a pear. Ripe quince fruits are hard and tart and seldom eaten raw, but are processed into marmalade, jam, paste (known as quince cheese) or alcoholic beverages.

The quince tree is grown as an ornamental plant for its attractive pale pink blossoms and other ornamental qualities.[1]

Description
Halved quince, with seeds and oxidation visible

The tree grows 5 to 8 metres (16 to 26 feet) high and 4 to 6 m (13 to 20 ft) wide. The fruit is 70 to 120 mm (3 to 4+1⁄2 in) long and 60 to 90 mm (2+1⁄2 to 3+1⁄2 in) across.

The immature fruit is green with dense grey-white fine hair, most of which rubs off before maturity in late autumn when the fruit changes colour to yellow with hard, strongly perfumed flesh. The leaves are alternately arranged, simple, 60–110 mm (2+1⁄2–4+1⁄2 in) long, with an entire margin and densely pubescent with fine white hairs. The flowers, produced in spring after the leaves, are white or pink, 50 mm (2 in) across, with five petals.

The seeds contain nitriles, which are common in the seeds of the rose family. In the stomach, enzymes or stomach acid or both cause some of the nitriles to be hydrolysed and produce hydrogen cyanide, which is a volatile gas. The seeds are only toxic if eaten in large quantities.[2]
Taxonomy

Four other species previously included in the genus Cydonia are now treated in separate genera. These are Pseudocydonia sinensis and the three flowering quinces of eastern Asia in the genus Chaenomeles. Another unrelated fruit, the bael, is sometimes called the "Bengal quince".

The modern name originated in the 14th century as a plural of quoyn, via Old French cooin from Latin cotoneum malum / cydonium malum, ultimately from Greek κυδώνιον μῆλον, kydonion melon "Kydonian apple".

Cydonia is included in the subfamily Amygdaloideae.[3]
Distribution and habitat
Commercially grown quince
Quince foliage and ripening fruit
Quince flowers
Quince nursery
Quince fruits in tree

Quince is native to the Hyrcanian forest region south of the Caspian Sea,[4][5] although it thrives in a variety of climates and can be grown successfully at latitudes as far north as Scotland. It should not be confused with its relatives, the Chinese quince, Pseudocydonia sinensis, or the flowering quinces of genus Chaenomeles, either of which is sometimes used as a culinary substitute.
History

The fruit was known to the Akkadians, who called it supurgillu; "quinces" (collective plural),[6] as well as in Judea of Israel during the Mishnaic era where it was called perishin (פרישין collective plural, or sing. prish);[7] quince flourished in the heat of the Mesopotamian plain, where apples did not. It was cultivated from an archaic period around the Mediterranean. Some ancients called the fruit "golden apples".[8]

The Greeks associated it with Cydonia on Crete, as the "Cydonian pome", and Theophrastus, in his Enquiry into Plants, noted that quince was one of many fruiting plants that do not come true from seed.[9] As a sacred emblem of Aphrodite, a quince figured in a lost poem of Callimachus that survives in a prose epitome: seeing his beloved in the courtyard of the temple of Aphrodite, Acontius plucks a quince from the "orchard of Aphrodite", inscribes its skin and furtively rolls it at the feet of her illiterate nurse, whose curiosity aroused, hands it to the girl to read aloud, and the girl finds herself saying "I swear by Aphrodite that I will marry Acontius". A vow thus spoken in the goddess's temenos cannot be broken.[10] Pliny the Elder mentions "numerous varieties" of quince in his Natural History and describes four.[11] The season of ripe quinces is brief: the Roman cookbook De re coquinaria of "Apicius" specifies in attempting to keep quinces, to select perfect unbruised fruits and keep stems and leaves intact, submerged in honey and reduced wine.[12]
Pests and diseases

Quince is used as a food plant by the larvae of some Lepidoptera species including brown-tail, Bucculatrix bechsteinella, Bucculatrix pomifoliella, Coleophora cerasivorella, Coleophora malivorella, green pug and winter moth.

While quince is a hardy shrub, it may develop fungal diseases in hot weather, resulting in premature leaf fall.[13] Quince leaf blight, caused by fungus Diplocarpon mespili, presents a threat in wet summers, causing severe leaf spotting and early defoliation, also affecting fruit to a lesser extent. It may also affect other Rosaceae plants such as hawthorn and medlar, but is typically less damaging than on quince.[14] Cedar-quince rust, caused by Gymnosporangium clavipes, requires two hosts to complete the fungal life cycle, one being a cedar (most commonly a juniper, Juniperus virginiana) and the other a rosacea. Appearing as red excrescence on various parts of the plant, it may affect quinces grown in vicinity of junipers.[15]
Cultivation

Quince is a hardy, drought-tolerant shrub which adapts to many soils of low to medium pH. It tolerates both shade and sun, but sunlight is required to produce larger flowers and ensure fruit ripening. It is a hardy plant that does not require much maintenance, and tolerates years without pruning or major insect and disease problems.[13] It is favored by landscape architects, such as Frederick Law Olmsted in the early 20th century, for its attractive blossoms.

Quince is cultivated on all continents in warm-temperate and temperate climates. It requires a cooler period of the year, with temperatures under 7 °C (45 °F), to flower properly. Propagation is done by cuttings or layering; the former method produces better plants, but they take longer to mature than by the latter. Named cultivars are propagated by cuttings or layers grafted on quince rootstock. Propagation by seed is not used commercially. Quince forms thick bushes, which must be pruned and reduced into a single stem to grow fruit-bearing trees for commercial use. The tree is self-pollinated, but it produces better yields when cross-pollinated.[13]

Fruits are typically left on the tree to ripen fully. In warmer climates, it may become soft to the point of being edible, but additional ripening may be required in cooler climates. They are harvested in late autumn, before first frosts.[13]

Quince is also used as rootstock for certain pear cultivars.[13] The resultant chimera is called + Pirocydonia danielii.

In Europe, quinces are commonly grown in central and southern areas where the summers are sufficiently hot for the fruit to fully ripen. They are not grown in large amounts; typically one or two quince trees are grown in a mixed orchard with several apples and other fruit trees. In the 18th-century New England colonies, for example, there was always a quince at the lower corner of the vegetable garden, Ann Leighton notes in records of Portsmouth, New Hampshire and Newburyport, Massachusetts.[16]: 243  Charlemagne directed that quinces be planted in well-stocked orchards. Quinces in England are first recorded in about 1275, when Edward I had some planted at the Tower of London.[17]
Cultivars

'Aromatnaya'
'Bereczki'
'Champion'
'Cooke's Jumbo' (syn. 'Jumbo')
'Dwarf Orange'
'Gamboa'
'Iranian'
'kashmiri Bumm tchoont'
'Isfahan'

'Le Bourgeaut'
'Lescovacz'
'Ludovic'
'Maliformis'
'Meeches Prolific'
'Morava'
'Orange' (syn. 'Apple quince')
'Perfume'

'Pineapple'
'Portugal' (syn. 'Lusitanica')
'Shams'
'Siebosa'
'Smyrna'
'Van Deman'
'Vranja' (syn. 'Bereczki')[18]

Quince production – 2020
Country	tonnes
Turkey	189,251
China	111,982
Uzbekistan	96,242
Iran	87,799
Morocco	57,700
Azerbaijan	39,365
World	696,861
Source: UN FAOSTAT[19]

The cultivar 'Vranja' Nenadovic has gained the Royal Horticultural Society's Award of Garden Merit.[20]
Production

In 2020 world production of quinces amounted to 696,861 tonnes, with Turkey and China growing a combined 43% of the world total (table).
Uses
Quinces, raw

Nutritional value per 100 g (3.5 oz)
Energy	238 kJ (57 kcal)
Carbohydrates	15.3 g
Dietary fibre	1.9 g
Fat	0.1 g
Protein	0.4 g
Vitamins	Quantity %DV†
Thiamine (B1)	2% 0.02 mg
Riboflavin (B2)	3% 0.03 mg
Niacin (B3)	1% 0.2 mg
Pantothenic acid (B5)	2% 0.081 mg
Vitamin B6	3% 0.04 mg
Folate (B9)	1% 3 μg
Vitamin C	18% 15 mg
Minerals	Quantity %DV†
Calcium	1% 11 mg
Iron	5% 0.7 mg
Magnesium	2% 8 mg
Phosphorus	2% 17 mg
Potassium	4% 197 mg
Sodium	0% 4 mg
Zinc	0% 0.04 mg
Other constituents	Quantity
Water	84 g
Link to USDA Database entry
Units μg = micrograms • mg = milligrams IU = International units
†Percentages are roughly approximated using US recommendations for adults. Source: USDA FoodData Central

Quinces are appreciated for their intense aroma, flavour, and tartness. However, most varieties of quince are too hard and tart to be eaten raw; even ripe fruits should be subjected to bletting by frost or decay to be suitable for consumption. However, they may be cooked or roasted and used for jams, marmalade, jellies, or pudding.[13]
As food

Some varieties of quince, such as 'Aromatnaya' and 'Kuganskaya' do not require cooking and can be eaten raw.[21] However, most varieties of quince are too hard, astringent and sour to eat raw unless "bletted" (softened by frost and subsequent decay).[22] High in pectin, they are used to make jam, jelly and quince pudding, or they may be peeled, then roasted, baked or stewed; pectin levels diminish as the fruit ripens.[23] The flesh of the fruit turns red after a long cooking with sugar by formation of anthocyanins.[24] The very strong perfume means they can be added in small quantities to apple pies and jam to enhance the flavor. Adding a diced quince to apple sauce will enhance the taste of the apple sauce with the chunks of relatively firm, tart quince. The term "marmalade", originally meaning a quince jam, derives from marmelo, the Portuguese word for this fruit.[25][26][27]
Quince cheese

Quince cheese is firm, sticky, sweet reddish hard paste made from the quince fruit, and originating from the Iberian peninsula. It is known as marmelada across the Portuguese-speaking world and as carne de membrillo or dulce de membrillo across the Spanish-speaking world, where it is used in a variety of recipes, eaten in sandwiches and with cheese, traditionally manchego cheese, or accompanying fresh curds. In Chile, boiled quince is popular in desserts such as the murta con membrillo that combines Chilean guava with quince.
Nutrition

A raw quince is 84% water, 15% carbohydrates, and contains negligible fat and protein (table). In a 100-gram (3+1⁄2-ounce) reference amount, the fruit provides 238 kilojoules (57 kilocalories) of food energy and a moderate amount of vitamin C (18% of the Daily Value), but no other micronutrients of significant quantity.
Alcoholic drink

In the Balkans and elsewhere, quince eau-de-vie (rakija) is made. For a quince rakija, ripe fruits of sweeter varieties are washed and cleared from rot and seeds, then crushed or minced, mixed with cold or boiling sweetened water and yeast, and left for several weeks to ferment. The fermented mash is distilled once, obtaining a 20–30 ABV, or twice, producing an approximately 60% ABV liquor. The two distillates may be mixed or diluted with distilled water to obtain the final product, containing 42–43% ABV.[28][29] Traditionally, it is not aged in wooden casks.

In the Alsace region of France and the Valais region of Switzerland, liqueur de coing made from quince is used as a digestif.

In Carolina in 1709, John Lawson allowed that he was "not a fair judge of the different sorts of Quinces, which they call Brunswick, Portugal and Barbary", but he noted "of this fruit they make a wine or liquor which they call Quince-Drink, and which I approve of beyond any that their country affords, though a great deal of cider and perry is there made, The Quince-Drink most commonly purges."[30]
Ornamental

Quince is one of the most popular species for deciduous bonsai specimens,[13] along with related Chinese quince and Japanese quince, native to Eastern Asia.
Medicinal value

According to research, quince helps reduce symptoms of early pregnancy such as nausea and vomiting.[31] Traditionally, the fruit has been used to cure digestive disorders; in fact, it promises to inhibit growth of H. Pylori, an ulcer causing bacteria in the stomach.[32]
Cultural associations

In "El licenciado Vidriera" by Miguel de Cervantes, the protagonist develops the delusion that he is made of glass after he eats a poisoned quince.
In Turkey, the expression ayvayı yemek (literally "to eat the quince") is used as a derogatory term indicating any unpleasant situation or a malevolent incident to avoid. This usage is likened to the rather bitter aftertaste of a quince fruit inside the mouth.
When a baby is born in the Balkans, a quince tree is planted as a symbol of fertility, love and life.[13]
Ancient Greek poets (Ibycus and Aristophanes, for example) used quinces (kydonia) as a mildly ribald term for teenage breasts.
In Plutarch's Lives, Solon is said to have decreed that "bride and bridegroom shall be shut into a chamber, and eat a quince together."[33]
In The Simpsons episode "Who Shot Mr. Burns? (Part One)", Homer sends Mr. Burns a box of chocolates with a family photo at the bottom. Burns and Smithers eschew the sour quince log, leaving Homer's face obscured in the photo.
The Goblin Market, a 19th century narrative poem by English poet, Christina Rossetti, references Quinces in its opening lines, which list myriad fruits.

Gallery

Still-life of quinces by Vincent van Gogh (1887), Albertinum, Dresden

Closeup of Russian 'Aromatnaya' quinces

Ripe Constantinople apple quinces from the Vogelsberg

See also

List of culinary fruits

References

Postman, Joseph (2009). "Cydonia oblonga: The unappreciated quince". Arnoldia. 67 (1): 2–9. JSTOR 42955445.
"Cydonia oblonga Quince PFAF Plant Database". pfaf.org.
Potter, D., et al. (2007). Phylogeny and classification of Rosaceae. Plant Systematics and Evolution. 266(1–2): 5–43. [Referring to the subfamily by the name "Spiraeoideae"]
Zohary, Daniel; Hopf, Maria; Weiss, Ehud (2012). Domestication of Plants in the Old World: The Origin and Spread of Domesticated Plants in Southwest Asia, Europe, and the Mediterranean Basin. OUP Oxford. p. 144. ISBN 978-0-19-954906-1.
Brickell, Christopher (2008). A-Z encyclopedia of garden plants. London: Dorling Kindersley. p. 1136. ISBN 978-1405332965.
Lauffenburger, Olivier (2006). "supurgillu". The Hittite Grammar Homepage, Akkadian Dictionary – via premiumwanadoo.com.
Maimonides (1967). "1:6". Mishnah commentary to tractate Uksin (original Judeo-Arabic אלספרג׳ל (Mossad HaRav Kook ed.). Jerusalem.
Lyle, Katie Letcher (2010) [first published 2004]. The Complete Guide to Edible Wild Plants, Mushrooms, Fruits, and Nuts: How to Find, Identify, and Cook Them (2nd ed.). Guilford, Conn.: FalconGuides. p. 110. ISBN 978-1-59921-887-8. OCLC 560560606.
Theophrastus. "Quince produces wild quince". Enquiry.
Skinner, Marilyn B. (2003). "Carmina Battiadae". Catullus in Verona: a Reading of the Elegiac Libellus. Columbus: Ohio State University Press. pp. 15ff. ISBN 978-0-8142-0937-0.
Pliny the Elder. Natural History. pp. xv.10.11.
Lacus Curtius. Apicius.
Carlton, Deb (2013). "Quince". In Cumo, Christopher (ed.). Encyclopedia of Cultivated Plants: From Acacia to Zinnia, Volume III. Santa Barbara, Calif.: ABC-CLIO. pp. 855–858. ISBN 978-1-59884-775-8.
"Quince leaf blight". Royal Horticultural Society. 2016. Retrieved 22 November 2016.
"Cedar-Quince Rust". RMissouri Botanical Garden. 2016. Retrieved 22 November 2016.
Leighton, Ann (1986) [first published 1976]. American Gardens in the Eighteenth Century: "For Use Or for Delight". Amherst: University of Massachusetts Press. ISBN 978-0-8702-3531-3.
Fearnley-Whittingstall, Hugh (7 October 2011). "Quince recipes". The Guardian.
"Agroforestry news quince cydonia oblonga". agroforestry.co.uk. Archived from the original on 6 October 2006.
"Quince production in 2017; Crops/World regions/Production total from picklists". UN Food and Agriculture Organization Corporate Statistical Database. 2018. Retrieved 19 May 2019.
"Cydonia oblonga 'Vranja' Nenadovic". Royal Horticultural Society. Retrieved 22 July 2013.
Online Database: National Germplasm Resources Laboratory. Beltsville, Maryland: USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network (GRIN). Retrieved 20 February 2011.
"Quince". herbs2000.com.
Alexander, S. (2004). The cook's companion. Penguin Australia. p. 609. ISBN 978-1-9209-8900-2.
McGee, H. (2004). On Food and Cooking. Hodder & Stoughton. p. 357. ISBN 978-0-684-80001-1.
Wilson, C. Anne (1999) [first published 1985]. The Book of Marmalade: Its Antecedents, Its History and Its Role in the World Today (Together with a Collection of Recipes for Marmalades and Marmalade Cookery) (Revised ed.). Philadelphia: University of Pennsylvania Press. ISBN 0-8122-1727-6.
Harper, Douglas (2001). "Marmalade". Online Etymology Dictionary – via Dictionary.com.
Ferraz, Rafaela (5 December 2018). "Why Portugal's Marmelada Tastes Nothing Like Marmalade". Gastro Obscura. Atlas Obscura. Retrieved 8 December 2018.
"Dunjevača izuzetne arome i ukusa" [Quince brandy of exceptional aroma and taste] (in Serbian). Poljoprivreda.info. 22 November 2003. Retrieved 3 February 2020.
"Kako da napravite kvalitetnu dunjevaču" [How to make a quality quince] (in Serbian). Agropress. 11 October 2015. Retrieved 3 February 2020.
Lawson, John (1709). A New Voyage to Carolina. Quoted in Leighton (1986), p. 243
Jafari-Dehkordi, E.; Hashem-Dabaghian, F.; Aliasl, F.; Aliasl, J.; Taghavi-Shirazi, M.; Sadeghpour, O.; Sohrabvand, F.; Minaei, B.; Ghods, R. (20 June 2017). "Comparison of quince with vitamin B6 for treatment of nausea and vomiting in pregnancy: a randomised clinical trial". Journal of Obstetrics and Gynaecology : The Journal of the Institute of Obstetrics and Gynaecology. 37 (8): 1048–1052. doi:10.1080/01443615.2017.1322046. PMID 28631509. S2CID 3673722.
Ashraf, M. U.; Muhammad, G.; Hussain, M. A.; Bukhari, S. N. (21 June 2016). "Cydonia oblonga M., A Medicinal Plant Rich in Phytonutrients for Pharmaceuticals". Frontiers in Pharmacology. 7: 163. doi:10.3389/fphar.2016.00163. PMC 4914572. PMID 27445806.
Wikisource: Lives by Plutarch, translated by John Dryden: Solon