Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
OrdoTherapsida
Cladus: Theriodontia
Subordo: Cynodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohort: Theria
Cohort: Eutheria
Cohort: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Ordo: Artiodactyla
Subordo: Whippomorpha
Infraordo: Cetacea
Cladus: †Archaeoceti
Familia: †Protocetidae
Genus: †Qaisracetus
Species: †Q. arifi
Name
Qaisracetus Gingerich, ul-Haq, Khan & Zalmout, 2001: 295
Type species: Qaisracetus arifi [Gingerich et al., 2001: 295]
References
Primary references
Gingerich, P.D., ul-Haq, M., Khan, I.H. & Zalmout, I.S. 2001. Eocene stratigraphy and archeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sumalian Range, Balochistan (Pakistan). Contributions from the Museum of Paleontology, The University of Michigan 30(11): 269–319. hdl: 2027.42/48661 Reference page.
Links
Qaisracetus – Taxon details on Fossilworks.
Qaisracetus is an extinct protocetid early whale known from the Eocene (Lutetian, 48.6 to 40.4 million years ago) of Baluchistan, Pakistan (31.0°N 70.2°E, paleocoordinates 12.4°N 65.7°E).[1][2]
Etymology
The genus is named after the Qaisrani Baloch tribe which assisted Gingerich and his team during their field work.[3] "Qaisra" is also etymologically close to the royal title used in Persian and many Indo-European languages (e.g. Kaiser, Czar, Caesar).[3] The species is named for Muhammad Arif, former paleontologist at the Geological Survey of Pakistan who contributed significantly to archaeocete paleontology in Pakistan.[4]
Description
Qaisracetus is known from a dozen specimens, all found in or near the type locality. Among them are several well-preserved elements, including a well-preserved skull, partial skulls and braincases, several vertebrae including an almost complete sacrum, a left innominate, ribs, and partial limb elements.[5]
Qaisracetus is smaller than Pappocetus and Babiacetus but larger than Indocetus.[6] Qaisracetus arifi is almost as complete as Rodhocetus kasranii, the most complete articulated skeleton of a protocetid, and they were similar in size: the latter had an estimated body weight of 620 kg (1,370 lb), compared to 590 kg (1,300 lb) for the former.[7]
Qaisracetus has a generalized protocetid skull with the external nares located relatively anteriorly (above C1) and a relatively broad frontal shield. The rostrum is more narrow in Qaisracetus than in Takracetus.[6]
The four sacral vertebrae are only partially fused: the first two are solidly fused, the third is locked in place by rib-like processes (a pleurapophyseal synchondrosis), while the fourth has a caudal (tail-like) morphology including two ventral chevron processes. The fusion between S1 and S2 distinguished Qaisracetus from other protocetids such as Protocetus, Rodhocetus, Gaviacetus, Natchitochia, and Georgiacetus. Qaisracetus' vertebrae are not dense and thick like in Eocetus.[8] In contrast to Qaisracetus, Rodhocetus has a sacrum where non of the vertebrae have fused centra, which is derived to be a protocetid, but Rodhocetus is primitive in retaining pleurapophyseal connections between all sacral vertebrae. The sacral morphology of Rodhocetus and Qaisracetus indicate protocetids represent a wide range of specializations, although which is ancestral to later whales is unclear.[7]
Notes and references
Qaisracetus in the Paleobiology Database. Retrieved July 2013.
Ander Dabh Shumali (Eocene of Pakistan) in the Paleobiology Database. Retrieved July 2013.
Gingerich et al. 2001, Etymology, p. 296
Gingerich et al. 2001, Etymology, p. 298
Gingerich et al. 2001, Hypodigm, p. 297
Gingerich et al. 2001, Diagnosis, p. 296
Gingerich et al. 2001, Discussion, pp. 307–8
Gingerich et al. 2001, Diagnosis, p. 296, 303, Fig. 23
Bibliography
Gingerich, Philip D; Haq, U; Khan, H; Zalmout, S (2001). "Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman range, Balochistan (Pakistan)". Contributions from the Museum of Paleontology, University of Michigan. 30 (11): 269–319. hdl:2027.42/48661. OCLC 50061585.
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