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Saxicola rubicola

Saxicola rubicola (*)

Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Aves
Subclassis: Carinatae
Infraclassis: Neornithes
Parvclassis: Neognathae
Ordo: Passeriformes
Subordo: Passeri
Parvordo: Passerida
Superfamilia: Muscicapoidea
Familia: Muscicapidae
Genus: Saxicola
Species: Saxicola rubicola
Subspecies: S. r. albofasciata - S. r. hibernans - S. r. rubicola

Name

Saxicola rubicola (Linnaeus, 1766)

The European Stonechat Saxicola rubicola (syn. S. torquatus rubicola) is a small passerine bird that was formerly classed as a subspecies of the Common Stonechat. Long considered a member of the thrush family Turdidae, genetic evidence has placed it and its relatives in the Old World flycatcher family Muscicapidae.

It is 11.5–13 cm long and weighs 13–17 g, slightly smaller than the European Robin. Both sexes have distinctively short wings, shorter than those of the more migratory Whinchat and Siberian Stonechat. The summer male has black upperparts, a black head, an orange throat and breast, and a white belly and vent. It also has white half-collar on the sides of its neck, a small white scapular patch on the wings, and a very small white patch on the rump often streaked with black. The female has brown upperparts and head, and no white neck patches, rump or belly, these areas being streaked dark brown on paler brown, the only white being the scapular patch on the wings and even this often being buffy-white.[1]

European Stonechats breed in heathland, coastal dunes and rough grassland with scattered small shrubs and bramble, open gorse, tussocks or heather. They are short-distance migrants or non-migratory, with part of the population (particularly from northeastern parts of the range, where winters are colder) moving south to winter further south in Europe and more widely in north Africa.[1]

The male's song is high and twittering like a Dunnock. Both sexes have a clicking call like stones knocking together.

Taxonomy

Two weakly defined subspecies are currently accepted:[1]

* Saxicola rubicola rubicola. In the south and east of its range, from Denmark southwest to Spain and northern Morocco, east to Poland and Ukraine, and southeast to Turkey.
* Saxicola rubicola hibernans. Northwestern Europe in Atlantic coastal areas, in southwestern Norway, Great Britain, Ireland and northwestern France. Birds in coastal Portugal are also often listed as this subspecies[1] but this is disputed.[2]

The two subspecies differ in colour intensity following Gloger's rule, with S. r. rubicola paler and with larger white patches in the drier European continental and mediterranean climates, and S. r. hibernans darker brown with less white in the humid Atlantic oceanic climate. They intergrade broadly where their ranges meet, from southeastern England[3] south through France and Spain, and many individuals are not identifiable to subspecies. Extreme examples of S. r. rubicola from the driest southern areas of its range such as the Algarve and Sicily are particularly pale and with a large white rump, and can be very similar to Siberian Stonechats in appearance.[2][4] nDNA microsatellite fingerprinting reveals a very small degree of separation between the two subspecies.[5]

In the past, the European Stonechat was generally considered conspecific with the Siberian Stonechat and African Stonechat, lumped together as Common Stonechat S. torquatus. A new review adding mtDNA cytochrome b sequence and nuclear DNA microsatellite fingerprinting evidence strongly supports their separation into distinct species.[1][5] Due to a misunderstanding of the rules of Zoological nomenclature, for a short time the name S. torquatus was erroneously used for the European Stonechat rather than the African Stonechat.[5]

Together with the Siberian and Fuerteventura Stonechats it constitutes eastern and western representatives of an Eurasian lineage; the Asian and European populations separated during the Late Pliocene or Early Pleistocene, roughly 1.5-2.5 mya, and Fuerteventura was colonised by western European or northwest African birds somewhat later in the Early Pleistocene, about 1-2 mya.[5]

The etymology of the English name derives from its call, sounding like two stones knocked together. The scientific name Saxicola means "rock-dweller", from Latin saxum, a rock + incola, dwelling in; and rubicola, "bramble-dweller", from Latin Rubus, brambles + incola; the subspecies name hibernans refers to Ireland (Latin, Hibernia).

References

1. ^ a b c d e Urquhart, E., & Bowley, A. (2002): Stonechats. A Guide to the Genus Saxicola. Christopher Helm, London. ISBN 0-7136-6024-4
2. ^ a b Siddle, J. P. (2006). Which subspecies of Common Stonechat breeds in coastal Portugal? British Birds 99: 372-374
3. ^ Kehoe, C. (2006). Racial identification and assessment in Britain: a report from the RIACT subcommittee. British Birds 99: 619-645 [Stonechat, pp. 635-636].
4. ^ Corso, A. (2001). Plumages of Common Stonechats in Sicily and comparison with vagrant Siberian Stonechats. British Birds 94: 315-318.
5. ^ a b c d Wink, M.; Sauer-Gürth, H. & Gwinner, E. (2002): Evolutionary relationships of stonechats and related species inferred from mitochondrial-DNA sequences and genomic fingerprinting. British Birds 95: 349-355. PDF fulltext

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