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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Cladus: Columbimorphae
Ordo: Columbiformes

Familia: Columbidae
Subfamilia: Columbinae
Genus: Pampusana
Species: P. beccarii – P. canifrons – P. erythroptera – P. hoedtii – P. jobiensis – P. kubaryi – P. rubescens – P. sanctaecrucis – P. stairi – P. xanthonura

Species extinctae: †P. ferruginea – †P. norfolkensis – †P. salamonis
Name

Pampusana Bonaparte, 1855
References

Bonaparte,C-L, 1855. Coup d'oeil sur l'ordre des pigeons Title page

Pampusana is a mid-sized genus of ground-dwelling doves (family Columbidae) which occur in rainforests in the Pacific region. They are not closely related to the American ground doves (genus Columbina and related genera).

Systematics and extinctions

This genus was previously named Alopecoenas Sharpe, 1899, but Pampusana Bonaparte, 1855 has priority.[1][2] Pampusana might be ranked as a (very small) subfamily, but the available data suggests that they are better considered part of a quite basal radiation of Columbidae which consists of many small and often bizarre lineages; for example, Goura and Otidiphaps which are ecologically convergent to Galliformes, and maybe even the famous didines (Raphinae). Currently about 10 species of Pampusana are extant. Of the larger genera of Columbidae, Pampusana is the one most affected by extinction: 3–4 species have disappeared since the 18th century, and most of the remaining are decreasing in numbers, threatened with extinction or have lost subspecies due to habitat destruction, invasive species, or overhunting. In addition, there are several species never studied alive but known from subfossil bones. These became extinct during prehistory of the Southern Pacific region (c. 13000 BC – 1400 BC).

The bones of Pampusana are distinct enough to usually recognize this genus with ease. Still, the evolutionary affiliation of extinct species is often uncertain. A considerable number of recognizably distinct now-extinct Polynesian taxa used to inhabit the area where the white-headed lineage (P. jobiensis, P. erythroptera) and the Melanesian expansion that brought about P. sanctaecrucis and P. stairi (and possibly P. rubescens) would have met.

These ground-doves were removed from Gallicolumba (which was non-monophyletic) and reassigned here.[2][3][4]
List of species

The genus contains 13 species of which 3 became extinct in historical times:[2]

Wetar ground dove, Pampusana hoedtii
White-breasted ground dove, Pampusana jobiensis
White-fronted ground dove, Pampusana kubaryi
Polynesian ground dove, Pampusana erythroptera
White-throated ground dove, Pampusana xanthonura
† Norfolk ground dove, Pampusana norfolkensis - extinct (c.1800)
Tongan ground dove, Pampusana stairi
Santa Cruz ground dove, Pampusana sanctaecrucis
† Tanna ground dove, Pampusana ferruginea - extinct (late 18th-19th century)
† Thick-billed ground dove, Pampusana salamonis - extinct (mid-20th century)
Marquesan ground dove, Pampusana rubescens
Bronze ground dove, Pampusana beccarii
Palau ground dove, Pampusana canifrons

A further 4 species became extinct in prehistoric times:

† Rota ground dove, Gallicolumba sp. - prehistoric
† Great ground dove, Pampusana nui - prehistoric
† Henderson ground dove, Pampusana leonpascoi - prehistoric
† New Caledonian ground dove, Pampusana longitarsus - prehistoric

References

Bruce, M.; Bahr, N.; David, N. (2016). "Pampusanna vs. Pampusana: a nomenclatural conundrum resolved, along with associated errors and oversights" (PDF). Bulletin of the British Ornithologists' Club. 136: 86–100.
Gill, Frank; Donsker, David, eds. (2019). "Pigeons". World Bird List Version 9.2. International Ornithologists' Union. Retrieved 25 June 2019.
Jønsson, K.A.; Irestedt, M.; Bowie, R.C.K.; Christidis, L.; Fjeldså, J. (2011). "Systematics and biogeography of Indo-Pacific ground-doves". Molecular Phylogenetics and Evolution. 59 (2): 538–543. doi:10.1016/j.ympev.2011.01.007. PMID 21256968.
Moyle, R.G.; Jones, R.M.; Andersen, M.J. (2013). "A reconsideration of Gallicolumba (Aves: Columbidae) relationships using fresh source material reveals pseudogenes, chimeras, and a novel phylogenetic hypothesis". Molecular Phylogenetics and Evolution. 66 (3): 1060–1066. doi:10.1016/j.ympev.2012.11.024. PMID 23220516.

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