Lagonomegopidaea

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Chelicerata
Classis: Arachnida
Ordo: Araneae
Subordo: Opisthothelae
Infraordo: Araneomorphae
Taxon: Neocribellatae
Series: Entelegynae
Superfamilia: Palpimanoidea

Familia: †Lagonomegopidae
Subfamiliae: Lagonomegopinae
Genera: Albiburmops – Archaelagonops – Burlagonomegops – Cymbiolagonops – Grandoculus – Lagonoburmops – Lagonomegops – Lineaburmops – Myanlagonops – Parviburmops – Paxillomegops – Picturmegops – Planimegops – Soplaogonomegops – Spinomegops – Zarqagonomegops
Name

Lagonomegopidae Eskov & Wunderlich, 1995

Type genus:Lagonomegops Eskov & Wunderlich, 1995
Synonymy

Grandoculidae Penney, 2011. Synonymy in Wunderlich, 2012.

References
Primary references

Eskov, K.Y. & Wunderlich, J. 1995(1994). On the spiders from Taimyr ambers, Siberia, with the description of a new family and with general notes on the spiders from the Cretaceous resins (Arachnida: Araneae). Beiträge zur Araneologie 4: 95–107.

Additional references

Kaddumi, H.F. 2007. The amber of Jordan. The oldest prehistoric insects in fossilized resin. Third edition. Eternal River Museum of Natural History, Amman. 298 pp. Abstract.
Penney, D. 2004. Cretaceous Canadian amber spider and the palpimanoidean nature of lagonomegopids. Acta Palaeontologica Polonica 49: 579–584. PDF.
Penney, D. 2005. The fossil spider family Lagonomegopidae in Cretaceous ambers with descriptions of a new genus and species from Myanmar. Journal of arachnology 33: 439–444. PDF.
Penney, D. 2006. The oldest lagonomegopid spider, a new species in Lower Cretaceous amber from Álava, Spain. Geologica acta 4(3): 377–382. Abstract PDF.
Penney, D. & Selden, P.A. 2006. Assembling the Tree of Life—Phylogeny of Spiders: a review of the strictly fossil spider families. In: Deltshev, C.; Stoev, P. (eds.) European arachnology 2005. Acta zoologica bulgarica suppl. (1): 25–39. PDF.
Wunderlich, J. 2012. On the fossil Spider (Araneae) fauna in Cretaceous ambers, with descriptions of new taxa in amber from Myanmar (Burma) and Jordan, and on the relationships of the superfamily Leptonetoidea. Beiträge zur Araneologie 7: 157–232. Reference page.
Wunderlich, J. 2015. On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae). Beiträge zur Araneologie 9: 21–408. Reference page.
Wunderlich, J. 2017. New and rare fossil spiders (Araneae) in Mid Cretaceous amber from Myanmar (Burma), including the description of extinct families of the suborders Mesothelae and Opisthothelae, as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae. Beiträge zur Araneologie 10: 72–279. Reference page.

Links

ION

Lagonomegopidae is an extinct family of spiders known from the Cretaceous period. Members of the family are distinguished by a large pair of eyes, positioned on the anterolateral flanks of the carapace, with the rest of the eyes being small. They have generally been considered members of Palpimanoidea, but this has recently been questioned.[1] Members of the family are known from the late Early Cretaceous (Albian) to near the end of the Late Cretaceous (Campanian) of Eurasia, North America and the Middle East, which was then attached to Africa as part of Gondwana. They are generally assumed to have been free living hunters as opposed to web builders.[1]
Diagnosis

Per Guo and Selden, 2019[1]

Chelicera with several peg teeth on promargin; true teeth present or absent on retromargin. Carapace with a pair of large posterior median eyes situated on anterolateral corner, other eyes tiny. Endites subtriangular, directed across the labium, almost meeting at the midline. Trichobothria present on leg tibia and metatarsus. Three tarsal claws, unpaired claw hook-like. Six spinnerets. Female palpal tarsi lacking a claw.

For the meaning of technical words, refer to the Glossary of spider terms.
Relationships

A phylogenetic analysis conducted in 2021 found that Lagonomegopidae formed the sister group to extant Palpimanoidea.[2]
Paleobiology
Reconstruction of female lagonomegopid with egg sac

Langonomegopids preserved with associated egg sacs and spiderlings indicate that females likely laid egg sacs in nests or hollows, and the young may have lived with the mother for some time after hatching.[3]
Genera

†Albiburmops Wunderlich 2017[4]
†Albiburmops annulipes Wunderlich 2017 Burmese amber, Myanmar, Cenomanian
†Archaelagonops Wunderlich 2012[5]
†Archaelagonops alavensis Penney 2006[6] Spanish amber, Albian
†Archaelagonops propinquus Wunderlich 2015[7] Burmese amber, Myanmar, Cenomanian
†Archaelagonops salticoides Wunderlich 2012[5] Burmese amber, Myanmar, Cenomanian
†Archaelagonops scorsum Wunderlich 2015[7] Burmese amber, Myanmar, Cenomanian
†Cymbiolagonops Wunderlich 2015[7]
†Cymbiolagonops cymbiocalcar Wunderlich 2015[7] Burmese amber, Myanmar, Cenomanian
†Grandoculus Penney 2004[8]
†Grandoculus chemahawinensis Penney 2004[8] Canadian amber, Campanian
†Hiatomegops Guo, Selden and Ren, 2021[2]
†Hiatomegops spinalis Guo, Selden and Ren, 2021 Burmese amber, Myanmar, Cenomanian
†Jinjumegops Park et al. 2019[9]
†Jinjumegops dalingwateri Park et al. 2019[9] Jinju Formation, South Korea, Albian
†Koreamegops Park et al. 2019[9]
†Koreamegops samsiki Park et al. 2019[9] Jinju Formation, South Korea, Albian
†Lagonoburmops Wunderlich 2012[5]
†Lagonoburmops plumosus Wunderlich 2012[5] Burmese amber, Myanmar, Cenomanian
†Lagonomegops Eskov and Wunderlich 1995[10]
†Lagonomegops americanus Penney 2005[11] New Jersey amber, Turonian
†Lagonomegops cor Pérez-de la Fuente et al. 2013[12] Spanish amber, Albian
†Lagonomegops sukatchevae Eskov and Wunderlich 1995[10] Taimyr amber, Santonian
†Lagonomegops tuber Wunderlich 2015[7] Burmese amber, Myanmar, Cenomanian
†Lineaburmops Wunderlich 2015[7]
†Lineaburmops beigeli Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Lineaburmops hirsutipes Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Lineaburmops maculatus Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Myanlagonops Wunderlich 2012[5]
†Myanlagonops gracilipes Wunderlich 2012 Burmese amber, Myanmar, Cenomanian
†Odontomegops Guo and Selden 2019[1]
†Odontomegops titan Guo and Selden 2019 Burmese amber, Myanmar, Cenomanian
†Parviburmops Wunderlich 2015[7]
†Parviburmops bigibber Wunderlich 2017[4] Burmese amber, Myanmar, Cenomanian
†Parviburmops brevipalpus Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Paxillomegops Wunderlich 2015[7]
†Paxillomegops brevipes Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Paxillomegops cornutus Wunderlich 2017[4] Burmese amber, Myanmar, Cenomanian
†Paxillomegops longipes Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Picturmegops Wunderlich 2015[7]
†Picturmegops signatus Wunderlich 2015 Burmese amber, Myanmar, Cenomanian
†Planimegops Wunderlich 2017[4]
†Planimegops parvus Wunderlich 2017 Burmese amber, Myanmar, Cenomanian
†Scopomegops Guo, Selden and Ren, 2021[2]
†Scopomegops fax Guo, Selden and Ren, 2021 Burmese amber, Myanmar, Cenomanian
†Soplaogonomegops Pérez-de la Fuente et al. 2013[12]
†Soplaogonomegops unzuei Pérez-de la Fuente et al. 2013 Spanish amber, Albian
†Spinomegops Pérez-de la Fuente et al. 2013[12]
†Spinomegops aragonensis Pérez-de la Fuente et al. 2013 Spanish amber, Albian
†Spinomegops arcanus Pérez-de la Fuente et al. 2013 Spanish amber, Albian
†Zarqagonomegops Kaddumi 2007[13]
†Zarqagonomegops wunderlichi Kaddumi 2007 Jordanian amber, Albian

References

Guo, Xiangbo; Selden, Paul A.; Shih, Chungkun; Ren, Dong (February 2020). "Two new lagonomegopid spiders (Arachnida: Araneae) from the mid-Cretaceous of Northern Myanmar, with comments on the superfamilial placement of Lagonomegopidae". Cretaceous Research. 106: 104257. doi:10.1016/j.cretres.2019.104257. S2CID 204270349.
Guo, Xiangbo; Selden, Paul A; Ren, Dong (2021-06-16). "New specimens from Mid-Cretaceous Myanmar amber illuminate the phylogenetic placement of Lagonomegopidae (Arachnida: Araneae)". Zoological Journal of the Linnean Society. 195 (2): 399–416. doi:10.1093/zoolinnean/zlab027. ISSN 0024-4082.
Guo, Xiangbo; Selden, Paul A.; Ren, Dong (2021-09-29). "Maternal care in Mid-Cretaceous lagonomegopid spiders". Proceedings of the Royal Society B: Biological Sciences. 288 (1959): 20211279. doi:10.1098/rspb.2021.1279. ISSN 0962-8452. PMC 8441120. PMID 34521253.
J. Wunderlich. 2017. New and rare fossil spiders (Araneae) in mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae, as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae. Ten Papers on Fossil and Extant Spiders (Araneae). Beiträge zur Araneologie 10:72-279
J. Wunderlich. 2012. On the fossil spider (Araneae) fauna in Cretaeous ambers, with descriptions of new taxa from Myanmar (Burma) and Jordan, and on the relationships of the superfamily Leptonetoidea. Beiträge zur Araneologie 7:157-232
Penney, D. (2006-01-12). "The oldest lagonomegopid spider, a new species in Lower Cretaceous amber from Álava, Spain". Geologica Acta. 4 (3): 377–382. doi:10.1344/105.000000351. ISSN 1696-5728.
J. Wunderlich. 2015. On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae). Mesozoic Spiders (Araneae): Ancient Spider Faunas and Spider Evolution, Beiträge zur Araneologie 9:21-408
D. Penney. 2004. Cretaceous Canadian amber spider and the palpimanoidean nature of lagonomegopids. Acta Palaeontologica Polonica 49:579-584
Park, Tae-Yoon S.; Nam, Kye-Soo; Selden, Paul A. (2019-08-03). "A diverse new spider (Araneae) fauna from the Jinju Formation, Cretaceous (Albian) of Korea". Journal of Systematic Palaeontology. 17 (15): 1271–1297. doi:10.1080/14772019.2018.1525441. ISSN 1477-2019. S2CID 92137182.
K. Y. Eskov and J. Wunderlich. 1995. On the spiders from Taimyr ambers, Siberia, with the description of a new family and with general notes on the spiders from the Cretaceous resins (Arachnida: Araneae). Beiträge zur Araneologie 4:95-107
D. Penney. 2005. The fossil spider family Lagonomegopidae. The Journal of Arachnology 33:439-444
Fuente, Ricardo Pérez-de la; Saupe, Erin E.; Selden, Paul A. (2013-02-06). "New lagonomegopid spiders (Araneae: †Lagonomegopidae) from Early Cretaceous Spanish amber". Journal of Systematic Palaeontology: 1–23. doi:10.1080/14772019.2012.725679. ISSN 1477-2019. S2CID 55375767.
H. F. Kaddumi. 2007. Amber of Jordan. The Oldest Prehistoric Insects in Fossilized Resin 1-298