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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Cladus: Phytophaga
Superfamilia: Chrysomeloidea

Familia: Chrysomelidae
Subfamilia: Chrysomelinae
Tribus: Timarchini
Genus: Timarcha
Subgenera: T. (Americanotimarcha) – T. (Metallotimarcha) – T. (Timarcha)
Overview of species (Nearctic)

T. cerdo – T. intricata
Overview of species (Palearctic)

T. aitanae – T. angusta – T. apicicornis – T. apricaria – T. apuana – T. araneiformis – T. arragonica – T. aurichalcea – T. balearica – T. brachydera – T. brevipennis – T. calceata – T. chalcosoma – T. chloropus – T. corallipes – T. corinthia – T. cornuta – T. corpulenta – T. crassaticollis – T. cyanescens – T. cyrenaica – T. daillei – T. djelfaica – T. durmitorensis – T. elliptica – T. endora – T. espanoli – T. fallax – T. fracassii – T. freyi – T. geniculata – T. gibba – T. globipennis – T. gnaptoroides – T. goettingensis – T. gougeletii – T. granadensis – T. henonii – T. hummelii – T. immarginata – T. insignis – T. insparsa – T. intermedia – T. interstitialis – T. janthinipes – T. kiesenwetteri – T. lusitanica – T. mairei – T. marani – T. marginicollis – T. maritima – T. maroccana – T. melas – T. melica – T. metallica – T. montana – T. monticola – T. montsiensis – T. nicaeensis – T. nitidipes – T. oblongula – T. obsoleta – T. olivieri – T. opacicollis – T. peresa – T. pfefferi – T. pimelioides – T. piochardi – T. pontavicei – T. pratensis – T. prujai – T. punctella – T. punica – T. recticollis – T. reticulata – T. rugosa – T. rugulosa – T. sardea – T. scabripennis – T. scutellaris – T. sicelidis – T. sinuatocollis – T. sobrina – T. sphaeroptera – T. spuria – T. staneki – T. stepaneki – T. strangulata – T. strophium – T. sublaevis – T. taborskyi – T. tangeriana – T. tarsata – T. temperei – T. tenebricosa – T. tenuicornis – T. theryi – T. tingitana – T. tortosensis – T. trapezicollis – T. tunisea – T. turbida – T. validicornis – T. variolosa – T. ventricosa

In synonymy: T. asturiensis – T. perezii
[Source: Petitpierre & Anichtchenko (2018)]
Name

Timarcha Samouelle, 1819: 213

Type species: Chrysomela tenebricosa Fabricius, 1775, by subsequent designation by Chevrolat, 1843: 655.

Chrysomelidae - Timarcha tenebricosa

Timarcha tenebricosa

References
Primary references

Samouelle, G. 1819. The entomologists' useful compendium; or an introduction to the knowledge of British Insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linné, and modern methods of arranging the Classes Crustacea, Myriapoda, spiders, mites and insects, from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance and usual situations of near 3,000 species of British Insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp., 12 pls. BHL. Reference page.

Additional references

Gómez-Zurita, J. & Kippenberg, H. 2010. Tribe Timarchini Motschulsky, 1860. Pp. 437–443. In: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, Stenstrup, Denmark. 924 pp. ISBN 978-87-88757-84-2. Google Books. Reference page.
Löbl, I. & Smetana, A.. 2011. Errata to Volumes 1 to 6. Pp. 23–63. In: Löbl, I. & Smetana, A. (eds) Catalogue of Palaearctic Coleoptera. Volume 7. Curculionoidea I. Apollo Books, Stenstrup, Denmark. 373 pp. ISBN 978-87-88757-93-4. Google Books. [corrects authority and type species used in Gómez-Zurita & Kippenberg, 2010]
Petitpierre, E. & Anichtchenko, A. 2018. Endophallus structure: a promising tool for cryptic species identification in Timarcha Samouelle, 1819 (Coleoptera: Chrysomelidae: Chrysomelinae). Zootaxa 4446(3): 361–383. DOI: 10.11646/zootaxa.4446.3.4 Paywall Reference page.
Petitpierre, E. & Anichtchenko, A. 2022. New descriptions of endophalli of Timarcha Samuelson, 1819 (Coleoptera, Chrysomelidae). Zootaxa 5091(1): 191–196. DOI: 10.11646/zootaxa.5091.1.9. Paywall. ResearchGate Open access. Reference page.
Riley, E.G., Clark, S.M. & Seeno, T.N. 2003. Catalog of the leaf beetles of America North of Mexico (Coleoptera: Megalopididae, Orsodacnidae and Chrysomelidae, excluding Bruchinae). Coleopterists Society, Special Publication No. 1. Sacramento, CA. 290 pp. ISBN 978-0-9726087-1-8. PDF. Reference page.
Vela, J.M., Alonso-Zarazaga, M.Á. & Daccordi, M. 2020. The species of Timarcha Samouelle, 1819 described by Linnaeus (Coleoptera, Chrysomelidae). ZooKeys 986: 55–80. DOI: 10.3897/zookeys.986.57158 Open access. Reference page.
Vela, J.M. & Daccordi, M. 2021. Description of a new species of the genus Timarcha Samouelle, 1819 from Northeastern Spain (Coleoptera, Chrysomelidae). Boletín de la Asociación Española de Entomología 45(3–4): 293–298. Online (with PDF download). Reference page.
Vela, J.M. & Daccordi, M. 2022. A study of the genus Timarcha Samouelle, 1819: the species from the South of the Iberian Peninsula (Coleoptera, Chrysomelidae). Suplementos del Boletín de la Asociación Española de Entomología 3: 1–70. hdl: 10481/72692 Open access. Reference page.

Links

Subfamily Chrysomelinae (Chrysomelidae) - atlas of leaf beetles of Russia

Timarcha is a genus of leaf beetles in the family Chrysomelidae, with more than 100 described species in three subgenera. The most widely known species is T. tenebricosa, the bloody-nosed beetle. All species are black, wingless organisms. Timarcha are herbivorous species, living mostly on plants belonging to Rubiaceae and Plumbaginaceae, although a few can feed on Brassicaceae and Rosaceae. Timarcha is the only member of the tribe Timarchini.
Species

These 20 species belong to the genus Timarcha:

Timarcha affinis Laboissière, 1937 g
Timarcha cerdo Stål, 1863 i c g
Timarcha cornuta Bechyné, 1944 g
Timarcha cyanescens Fairmaire, 1862 g
Timarcha coarcticollis Fairmaire, 1873 g
Timarcha daillei Laboissière, 1939 g
Timarcha goettingensis (Linnaeus, 1758) g
Timarcha insparsa
Timarcha interstitialis Fairmaire, 1862 g
Timarcha intricata Haldeman, 1853 i c g b
Timarcha italica Herrich-Schäffer, 1838 g
Timarcha lugens
Timarcha marginicollis
Timarcha maritima Perris, 1855 g
Timarcha metallica (Laicharting, 1781) g
Timarcha monticola Dufour, 1851 g
Timarcha nicaeensis A.Villa & G.B.Villa, 1835 g
Timarcha obsoleta Laboissière, 1937 g
Timarcha recticollis Fairmaire, 1862 g
Timarcha sardea (Villa, 1835) g
Timarcha sinuatocollis Fairmaire, 1862 g
Timarcha strangulata Fairmaire, 1862 g
Timarcha temperei Jeanne, 1965 g
Timarcha tenebricosa (Fabricius, 1775) g (Bloody-nosed beetle)

Data sources: i = ITIS,[2] c = Catalogue of Life,[3] g = GBIF,[4] b = Bugguide.net[5]
References

Vela, J.M.; Alonso-Zarazaga, M.Á.; Daccordi, M. (2020). "The species of Timarcha Samouelle, 1819 described by Linnaeus (Coleoptera, Chrysomelidae)". ZooKeys (986): 55–80. doi:10.3897/zookeys.986.57158. PMC 7661484. PMID 33223880.
"Timarcha Report". Integrated Taxonomic Information System. Retrieved 2018-05-04.
"Browse Timarcha". Catalogue of Life. Retrieved 2018-05-04.
"Timarcha". GBIF. Retrieved 2018-05-04.
"Timarcha Genus: Information". BugGuide.net. Retrieved 2018-05-04.

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