Speyeria

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Heliconiinae
Tribus: Argynnini
Genus: Speyeria
Species: S. adiaste – S. aglaja – S. alexandra – S. aphrodite – S. atlantis – S. callippe – S. carolae – S. clara – S. coronis – S. cybele – S. diana – S. edwardsii – S. egleis – S. hesperis – S. hydaspe – S. idalia – S. mormonia – S. nokomis – S. zerene
Name

Speyeria Scudder, 1872

Type species: Papilio idalia Drury, [1773].
Synonyms

Semnopsyche Scudder, 1875
Mesoacidalia Reuss, 1926
Proacidalia Reuss, 1926
Neoacidalia Reuss, 1926

References

Campbell, E.O., Gage, E.V., Gage, R.V. & Sperling, F.A.H. 2020. Single nucleotide polymorphism‐based species phylogeny of greater fritillary butterflies (Lepidoptera: Nymphalidae: Speyeria) demonstrates widespread mitonuclear discordance. Systematic Entomology 45(2): 269–280. DOI: 10.1111/syen.12393 Reference page.
De Moya, R. S., Savage, W. K., Tenney, C., Bao, X., Wahlberg, N. & Hill, R. I., 2017. Interrelationships and diversification of Argynnis Fabricius and Speyeria Scudder butterflies. Systematic Entomology 42(4): 635–649. DOI: 10.1111/syen.12236 Reference page.
dos Passos, C.F. & Grey, L.P. 1945. Systematic catalog of Speyeria (Lepidoptera, Nymphalidae) with designations of types and fixations of type localities. American Museum Novitates 1297: 1–30. Reference page.
Dunford, J. C. 2009. Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae). Insecta Mundi 0090: 1–74. PDF Reference page.
Emmel, T. & Austin, G., 1998. What is Argynnis carolae? The Cytotaxonomy and Systematic Position of a Speyeria Relic from Nevada (Lepidoptera: Nymphalidae). In Systematics of Western North American Butterflies. 34: 443-450. Reference page.
Simonsen, T.J. 2006. Fritillary phylogeny, classification, and larval host plants: reconstructed mainly on the basis of male and female genitalic morphology (Lepidoptera: Nymphalidae: Argynnini). Biological Journal of the Linnean Society 89(4): 627–673. DOI: 10.1111/j.1095-8312.2006.00697.x Open access. Reference page.
Simonsen, T.J., Wahlberg, N., Brower, A.V.Z. & de Jong, R. 2006. Morphology, molecules and fritillaries: approaching a stable phylogeny for Argynnini (Lepidoptera: Nymphalidae). Insect Systematics & Evolution 37: 405–418. DOI: 10.1163/187631206788831407 Paywall. PDF. Reference page.

Speyeria, commonly known as greater fritillaries, is a genus of butterflies in the family Nymphalidae commonly found in North America, Europe, and Asia. Some authors used to consider this taxon a subgenus of Argynnis, but it has been reestablished as a separate genus in 2017.[1]

Species

The genus has 3 species in Eurasia (these were formerly known as genus Mesoacidalia, now a synonym of Speyeria):

Speyeria aglaja (Linnaeus, 1758) – Dark green fritillary
Speyeria alexandra (Ménétriés, 1832)
Speyeria clara (Blanchard, [1844])

and 16 species in North America:[1]

Speyeria diana (Cramer, [1777]) – Diana fritillary
Speyeria cybele (Fabricius, 1775) – great spangled fritillary
Speyeria aphrodite (Fabricius, 1787) – Aphrodite fritillary
Speyeria idalia (Drury, [1773]) – regal fritillary
Speyeria nokomis (Edwards, 1862) – Nokomis fritillary
Speyeria edwardsii (Reakirt, 1866) – Edward's fritillary
Speyeria coronis (Behr, 1864) – Coronis fritillary
Speyeria zerene (Boisduval, 1852) – zerene fritillary
Speyeria carolae (dos Passos & Grey, 1942) – Carole's fritillary
Speyeria callippe (Boisduval, 1852) – callippe fritillary
Speyeria egleis (Behr, 1863) – Egleis fritillary
Speyeria adiaste (Edwards, 1864) – unsilvered fritillary or Adiaste fritillary
Speyeria atlantis (Edwards, 1862) – Atlantis fritillary
Speyeria hesperis (Edwards, 1864) – northwestern fritillary
Speyeria hydaspe (Boisduval, 1869) – Hydaspe fritillary
Speyeria mormonia (Boisduval, 1869) – Mormon fritillary

References

Savela, Markku. "Speyeria Scudder, 1872". Lepidoptera and Some Other Life Forms. Retrieved October 1, 2019.

Wikimedia Commons has media related to Speyeria.
Wikispecies has information related to Speyeria.
Further reading
Glassberg, Jeffrey (2001). Butterflies through Binoculars: The West.
Guppy, Crispin S. and Shepard, Jon H. (2001). Butterflies of British Columbia.
James, David G. and Nunnallee, David (2011). Life Histories of Cascadia Butterflies.
Pelham, Jonathan (2008). Catalogue of the Butterflies of the United States and Canada.
Pyle, Robert Michael (2002). The Butterflies of Cascadia.
De Moya, Robert S.; Savage, Wesley K.; Tenney, Chris; Bao, Xiaoshan; Wahlberg, Niklas; Hill, Ryan I. (2017). "Interrelationships and diversification of Argynnis Fabricius and Speyeria Scudder butterflies". Systematic Entomology. 42 (4): 635–649. doi:10.1111/syen.12236.
Campbell, Erin O.; Gage, Edward V.; Gage, Randy V.; Sperling, Felix A. H. (2020). "Single nucleotide polymorphism‐based species phylogeny of greater fritillary butterflies (Lepidoptera: Nymphalidae: Speyeria) demonstrates widespread mitonuclear discordance". Systematic Entomology. 45 (2): 269–280. doi:10.1111/syen.12393.