Pentatominae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Heteroptera
Infraordo: Pentatomomorpha
Superfamilia: Pentatomoidea

Familia: Pentatomidae
Subfamilia: Pentatominae
Tribus: Aeliini - Aeptini - Aeschrocorini - Agaeini - Agonoscelidini - Amyntorini - Antestiini - Axiagastini - Bathycoeliini - Cappaeini - Carpocorini - Catacanthini - Caystrini - Chlorocorini - Coquereliini - Degonetini - Diemeniini - Diplostirini - Diploxyini - Eurysaspini - Eysarcorini - Halyini - Hoplistoderini - Lestonocorini - Mecideini - Memmiini - Menidini - Myrocheini - Nealeriini - Nezarini - Opsitomini - Pentamyrmecini - Pentatomini - Phricodini - Piezodorini - Procleticini - Rhynchocorini - Rolstoniellini - Sciocorini - Sephelini - Strachiini - Triplatygini - Incertae sedis

A

Abadia – Acanthidiellum – Accarana – Acesines – Acledra – Acoloba – Acrocorisellus – Acrosternum – Acrozangis – Actuarius – Acuticeps – Adelaidena – Adelolcus – Adevoplitus – Adria – Aednulus – Aednus – Aegaleus – Aeladria – Aelia – Aeliavuori – Aeliomorpha – Aeliopsis – Aeliosoma – Aenaria – Aeptus – Aeschrocoris – Aeschrus – Aesula – Aethemenes – Afrania – Afraniella – Agaeus – Agatharchoides – Agathocles – Aglaophon – Agonoscelis – Agroecus – Ahmadiana – Alcaeus – Alcimocoris – Alciphron – Aleixus – Allecbola – Alloeoglypta – Alphenor – Amasenus – Amatembuna – Amauromelpia – Ambiorix – Amblybelus – Amblycara – Ambohicorypha – Ameridalpa – Amirantea – Amphidexius – Amphimachus – Anaca – Anaxarchus – Anaxilaus – Anaximenes – Anchesmus – Anchises – Andocides – Anoano – Anolcus – Antestia – Antestiella – Antestiopsis – Antheminia – Apines – Aplerotus – Apodiphus – Araducta – Arniscus – Arocera – Artiazontes – Arvelius – Aspavia – Aspideurus – Asyla – Atelias – Atelocera – Auletrissa – Austromalaya – Auxentius – Avicenna – Axiagastus –
B

Bachesua – Bagrada – Banasa – Banya – Baracellus – Bathrus – Bathycoelia – Belopis – Benia – Berecynthus – Bergrothina – Biprorulus – Birketsmithia – Boea – Boerias – Bolaca – Bonacialis – Boocoris – Bourginia – Brachycoris – Brachymna – Brachynema – Brasilania – Brepholoxa – Brizica – Brizocoris – Brochymena – Bromocoris – Bucerocoris – Burma – Buthumka –
C

Cahara – Caonabo – Capivaccius – Capnoda – Cappaea – Caracia – Carbula – Carenoplistus – Caribo – Caridillus – Caridophthalmus – Carpocoris – Carvalhocoris – Catacanthus – Catalampusa – Caura – Cauromorpha – Caystrus – Cellobius – Cephaloplatus – Cervicoris – Chalazonotum – Chalcocoris – Chaubattiana – Chinavia – Chlorochroa – Chlorocoris – Chloropepla – Chraesus – Chroantha – Chrysodarecus – Cleoqueria – Cnemobia – Cnephosa – Coctoteris – Codophila – Coenomorpha – Coenus – Commius – Compsoprepes – Cooperocoris – Copeocoris – Coponia – Coquerelia – Coquerelidea – Corisseura – Cosmopepla – Cradia – Cratonotus – Cresphontes – Critheus – Curatia – Cuspicona – Cyptocephala –
D

Dabessus – Dalpada – Dardjilingia – Decellella – Degonetus – Delegorguella – Dendrites – Dendrocoris – Desertomenida – Diaphyta – Dichelops – Dictyometis – Dictyotus – Diemenia – Diplostira – Diploxys – Dippilana – Discimita – Disderia – Dissocolpus – Dollingiana – Dolycoris – Drinostia – Dryadantestia – Dryadocoris – Dunnius – Durmia – Dymantiscus – Dyroderes – Dysnoetus –
E

Ectenus – Eipeliella – Elanela – Elemana – Elsiella – Eludocoris – Empiesta – Ennius – Epipedus – Erachtheus – Eremophilacoris – Eribotes – Erlangerella – Erthesina – Eudolycoris – Eudryadocoris – Eupaleopada – Eurinome – Eurus – Eurydema – Eurymenida – Eurynannus – Eurysaspis – Euschistus – Everardia – Evoplitus – Exithemus – Eysarcoris –
F

Faizuda – Fecelia – Flaminia –
G

Gadarscama – Galedanta – Geomorpha – Gilippus – Glaucias – Glaucioides – Glottaspis – Glyphepomis – Goilalaka – Gomphocranum – Grazia – Grossiana – Grossimenia – Gulielmus – Gwea – Gynenica –
H

Halyabbas – Halycorypha – Halydicoris – Halyoides – Halyomorpha – Halys – Hegelochus – Heissocoris – Hermolaus – Hillieria – Himalayacoris – Hippotiscus – Hoffmanseggiella – Holcogaster – Holcostethus – Homalogonia – Hoplistodera – Humria – Hymenarcys – Hymenomaga – Hypanthracos – Hyparete – Hypatropis – Hypaulacus – Hypsithocus – Hyrmine –
I

Ilipla – Indrapura – Iphiarusa – Ippatha – Iskenderia – Izharocoris –
J

Janeirona – Jugalpada – Jitka –
K

Kahlamba – Kalkadoona – Kamaliana – Kapunda – Kaschmirocoris – Keleacoris – Kermana – Kilimacoris – Kitsoniocoris – Koogobatha – Kumbutha – Kurumana – Kyrtalus –
L

Ladeaschistus – Lagynotomus – Lakhonia – Laprius – Lathraedoeus – Lattinellica – Lattinidea – Lelia – Leovitius – Lerida – Leridella – Lestonocoris – Liicoris – Liodermion – Lobepomis – Lodosia – Lodosocoris – Lojus – Lokaia – Loxa – Lubentius – Luridocimex –
M

Macrocarenoides – Macrocarenus – Macromolus – Madates – Madecorypha – Manoriana – Marghita – Mariomella – Massocephalus – Mathiolus – Mayrinia – Mecidea – Mecocephala – Memmia – Menaccarus – Menecles – Menedemus – Menestheus – Menida – Menudo – Meridalpa – Meridindia – Mezessea – Millotia – Mimikana – Mimula – Mimulocoris – Modicia – Monteithiella – Mormidea – Mormidella – Morna – Moromorpha – Mulungua – Munduala – Munshiana – Murgantia – Muscanda – Mustha – Myappena – Mycoolona – Mygoodano – Myota – Myrochea –
N

Neagenor – Nealeria – Nene – Neoacrosternum – Neococalus – Neocoquerelidea – Neoderoploa – Neodius – Neodorpius – Neodymantis – Neogynenica – Neohalys – Neojurtina – Neolodosocoris – Neomazium – Neoplautia – Neoptolemus – Neostrachia – Neosurenus – Neotibilis – Neottiglossa – Nesagaeus – Nesobius – Nevisanus – Nezara – Niarius – Nimbocoris – Niphe – Nkolbissonicoris – Nocheta – Notius – Novatilla –
O

Obuducoris – Ochrophara – Ochrorrhaca – Ochyrotylus – Ocirrhoe – Odmalea – Oebalus – Oenopiella – Ogmocoris – Okeanos – Omyta – Oncinoproctus – Oncocoris – Oncotropis – Ooldeon – Opsitoma – Orthoschizops – Otantestia – Ouscha –
P

Padaeus – Pallantia – Palomena – Parabrochymena – Parachinavia – Paracritheus – Paradictyotus – Paragenor – Parahypatropis – Paralcimocoris – Paraleria – Paralerida – Paramecocephala – Paramecocoris – Paramenestheus – Paranevisanus – Parantestia – Parapoecilometis – Paratibilis – Paratibraca – Parentheca – Parocirrhoe – Parodius – Parodmalea – Parvacrena – Patanius – Paterculus – Pausias – Pedinonotus – Pegala – Pellaea – Peltasticus – Pentamyrmex – Pentatoma – Pentatomiana – Petalaspis – Phaeocoris – Phalaecus – Pharypia – Phavorinus – Phricodus – Piezodorus – Pirricoris – Placocoris – Placosternum – Platacantha – Platencha – Platycoris – Plautia – Pododus – Poecilometis – Poecilotoma – Polycarmes – Poriptus – Porphyroptera – Praetextatus – Pretorius – Priassus – Prionaca – Prionocompastes – Prionosoma – Prionotocoris – Procleticus – Proxys – Prytanicoris – Pseudaelia – Pseudapines – Pseudatelus – Pseudevoplitus – Pseudoacrosternum – Pseudobebaeus – Pseudolerida – Pseudoncocoris – Pugione –
R

Ramosiana – Rhaphigaster – Rhombocoris – Rhynchocoris – Rhyncholepta – Rhyssocephala – Riaziana – Riazocoris – Rideriana – Rio – Risbecella – Risibia – Roferta – Rolstoniellus – Rubiconia – Runibia –
S

Sabaeus – Saceseurus – Salixocoris – Saontarana – Sarju – Sciocoris – Sciomenida – Scribonia – Scylax – Seansonius – Senectius – Sennertus – Sephela – Sepontia – Sepontiella – Serdia – Sibaria – Similliserdia – Sinometis – Socantestia – Solenocoris – Solenogaster – Spermatodes – Spinalanx – Sronachlachar – Stachyomia – Stagonomus – Staria – Starioides – Steleocoris – Stenozygum – Stictochilus – Strachia – Stysiana – Stysicoris – Surenus –
T

Tachengia – Tantesia – Taurocerus – Taurodes – Tenerva – Tepa – Tepperocoris – Terania – Theloris – Theseus – Thestral – Tholosanus – Thoreyella – Thryptomenecoris – Thyanta – Tibilis – Tibraca – Tinganina – Tipulparra – Tisia – Tmetopis – Tolumnia – Trachyops – Trichopepla – Tricompastes – Trincavellius – Tripanda – Triplatyx – Trochiscocoris – Tropicorypha – Turrubulana – Tyomana – Tyoma –
U

Udonga – Umgababa – Utana – Utheria –
V

Vadonidea – Valescus – Veterna – Vidada – Vitellus – Vitruvius – Vulsirea –
X

Xiengia –
Z

Zangiola – Zaplutus – Zhengius – Zimmerana – Zouicoris
Name

Pentatominae Leach, 1815:121
References
Primary references

Leach, W.E. 1815. Entomology [pp. 57–172]. In: Brewster, D. (Ed). Brewster’s Edinburgh Encyclopedia. Volume IX [part I]. W. Blackwood, J. Waugh, etc., Edinburgh, 764 pp. BHL Reference page. [p. 121]

Additional references

Cioato, A., Bianchi, F.M., Eger, J. & Grazia, J. 2015. New species of Euschistus (Euschistus) from Jamaica, Euschistus (Mitripus) and Ladeaschistus from southern South America (Hemiptera: Heteroptera: Pentatomidae: Pentatominae). Zootaxa 4048(4): 565–574. DOI: 10.11646/zootaxa.4048.4.7. Preview (PDF) Reference page.
Faúndez, E.I.; Carvajal, M.A. 2011: Catalog of Chilean Pentatominae Leach, 1815 (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 2835: 53–60. Preview
Kment, P.; Jindra, Z.; Rider, D.A. 2014: New synonymies and new records of Afrotropical and Madagascan Pentatominae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa 3866(3): 371–397. DOI: 10.11646/zootaxa.3866.3.4 Reference page.
Kment, P. & Baena, M. 2015. A redescription of the endemic Madagascan genus Tricompastes (Hemiptera: Heteroptera: Pentatomidae). Zootaxa 4044(1): 65–78. DOI: 10.11646/zootaxa.4044.1.3. ResearchGate Reference page.
Rider, D.A., Zheng, L.Y. & Kerzhner, I.M. 2002. Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica rossica 11(1): 135–153. ResearchGate Open access. Reference page.
Rider, D.A. 2006. Family Pentatomidae. p. 233–403, In: Aukema, A. & Rieger, C. (eds). Catalogue of the Heteroptera of the Palaearctic Region, vol 5. The Netherlands Entomological Society, Amsterdam. ResearchGate Open access PDF. Reference page. [p. 246]
Rider, D.A., Schwertner, C.F., Davidová-Vilímová, J., Rédei, D., Kment, P. & Thomas, D.B. 2018. Higher systematics of the Pentatomoidea.. p. 25–204, In: McPherson, J.E. (ed). Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press. Taylor & Francis group. Boca Raton, FL. Reference page. [p. 60]
Salini, S. 2016. Redescription of Dardjilingia Yang (Hemiptera: Heteroptera: Pentatomidae) from India. Zootaxa 4144(1): 131–137. DOI: 10.11646/zootaxa.4144.1.9. Reference page.

Links

Pentatominae Leach, 1815 – Taxon details on Integrated Taxonomic Information System (ITIS).
Classification of Pentatomoidea. North Dakota State University[1].

Vernacular names
日本語: カメムシ亜科

Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera.[1] Species in this subfamily are phytophages and several of them are considered agricultural pests.[2] Some invasive pentatomines such as Halyomorpha halys and Bagrada hilaris have been considered household pests.[3] Higher systematics of the group have been revised by Rider et al.[4]
Tribes and selected genera

Tribes within this subfamily include:[5]

Aelini Douglas & Scott, 1865
Aelia Fabricius, 1803
Neottiglossa Kirby, 1837
Aeptini Stål, 1871
Aeschrocorini Distant, 1902
Agaeini Cachan, 1952
Agonoscelidini Atkinson, 1888
Amyntorini Distant, 1902
Antestiini Distant, 1902
Aulacetrini Mulsant & Rey, 1866
Axiagastini Atkinson, 1888
Bathycoeliini Atkinson, 1888
Cappaeini Atkinson, 1888
Halyomorpha Mayr, 1864
Tolumnia Stål, 1867
Adelolcus - Benia (insect) - Boerias - Cappaea - Caura - Cauromorpha - Erlangerella - Halycorypha - Halydicoris - Halyoides - Homalogonia - Hymenomaga - Lerida (insect) - Leridella - Lokaia - Massocephalus - Paralerida - Prytanicoris - Tenerva - Tripanda - Tropicorypha - Veterna
Carpocorini Mulsant & Rey, 1866
Antheminia Mulsant & Rey, 1866
Carpocoris Kolenati, 1846
Chlorochroa Stål, 1872
Dolycoris Mulsant & Rey, 1866
Graphorn Faúndez, Rider, & Carvajal, 2017
Peribalus Mulsant & Rey, 1866
Rubiconia Dohrn, 1860
Thestral Faúndez & Rider, 2014
Catacanthini Atkinson, 1888
Catacanthus Spinola, 1837
Caystrini Ahmad & Afzal, 1979
Chlorocorini
Coquereliini Cachan, 1952
Degonetini Azim & Shafee, 1984
Diemeniini Kirkaldy, 1909
Diplostirini Distant, 1902
Diploxyini Atkinson, 1888
Eurysaspidini Atkinson, 1888
Eysarcorini Mulsant & Rey, 1866
Carbula Stål, 1864
Eysarcoris Hahn, 1834
Halyini Amyot & Serville, 1843
Hoplistoderini Atkinson, 1888
Lestonocorini hmad & Mohammad, 1980
Mecideini Distant, 1902
Memmiini Cachan, 1952
Menidini Atkinson, 1888
Myrocheini Stål, 1871
Nealeriini Cachan, 1952
Nezarini Atkinson, 1888
Palomena Mulsant & Rey, 1866
Opsitomini Cachan, 1952
Pentamyrmexini
Pentamyrmex spinosus
Pentatomini Leach, 1815
Pentatoma Olivier, 1789
Phricodini Cachan, 1952
Piezodorini Atkinson, 1888
Rhynchocorini Stål, 1871
Rolstoniellini Rider, 1997
Sciocorini Amyot & Serville, 1843
Sciocoris Fallén, 1829
Strachiini Mulsant & Rey, 1866
Stenozygum coloratum (Klug, 1845)
Eurydema Laporte, 1833

Notable species include:

Acrostemum hilare – green stink bug
Bagrada hilaris – bagrada bug
Halyomorpha halys – brown marmorated stink bug
Pentatoma rufipes – forest bug
Piezodorus lituratus – gorse shield bug

References

Faúndez, E. I.; Rider, D. A.; Garcete-Barret, B. (2018). "New records of Alcaeorrhynchus grandis (Dallas, 1851)(Heteroptera: Pentatomidae) from Paraguay, with description of a teratological case". Boletín del Museo Nacional de Historia Natural del Paraguay. 22 (1): 27–29.
Faúndez, E. I.; Rider, D. A. (2017). "The brown marmorated stink bug Halyomorpha halys (Stål, 1855)(Heteroptera: Pentatomidae) in Chile". Arquivos Entomolóxicos. 17: 305–307.
Faúndez, E. I. (2018). "From agricultural to household pest: The case of the painted bug Bagrada hilaris (Burmeister)(Heteroptera: Pentatomidae) in Chile". Journal of Medical Entomology. 55 (5): 1365–1368. doi:10.1093/jme/tjy091. PMID 29931070.
Rider, D.A., Schwertner, C.F., Vilimová, J., Redei, D., Kment, P. & D.B. Thomas. 2018. Pp. 25-204. Higher systematics of Pentatomidea in McPherson, J. E. (Ed.). Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Florida.
Zicha, Ondrej. "BioLib: Biological library". www.biolib.cz.