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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Staphylinoidea

Familia: Staphylinidae
Subfamilia: Aleocharinae
Tribus: Myllaenini
Genus: Bryothinusa
Species: B. algarum – B. cameroni – B. catalinae – B. celebensis – B. chani – B. chengae – B. fluenta – B. gangjinensis – B. grootaerti – B. hauseri – B. hongkongensis – B. japonica – B. koreana – B. madecassa – B. minuta – B. nakanei – B. neoguineensis – B. orousseti – B. papuensis – B. parvula – B. perexilis – B. peyerimhoffi – B. rothi – B. sakishimana – B. samoensis – B. sawadai – B. sinensis – B. subtilissima – B. testacea – B. testaceipennis – B. tsutsuii
Name

Bryothinusa Casey, 1904: 312

Type species: Bryothinusa catalinae Cameron, 1904, fixed by original designation.

Synonymy

Anopsisus Bernhauer, 1929: 187
Type species: Phytosus microphthalmus Bernhauer, 1929 (= Diglossa cameroni Fauvel, 1904), fixed by monotypy.
Halesthenus Sawada, 1955: 83 [new synonym in Sawada, 1971: 81]
Type species: Halesthenus minutus Sawada, 1955, fixed by original designation.

References
Primary references

Casey, T.L., 1904: On some new Coleoptera, including five new genera. Canadian entomologist, 36: 312–323. [original description: p. 312]
Sawada, K. 1971. Aleocharinae (Staphylinidae, Coleoptera) from the intertidal zone of Japan. Publications of the Seto Marine Biological Laboratory 19(2–3): 81–110. Handle Reference page. [syn. nov.: p. 81]

Additional references

Jeon, M.-J.; Ahn, K.-J. 2009: Description of late-instars of Bryothinusa koreana Ahn and Jeon (Coleoptera: Staphylinidae: Aleocharinae) by association of life stage based on DNA sequence data. Florida entomologist, 92: 367–373. DOI: 10.1653/024.092.0224
Liu, T-T., Ono, H. & Maruyama, M. 2020. A new species and new record of the intertidal rove beetle genus Bryothinusa Casey (Staphylinidae: Aleocharinae: Myllaenini) from Japan. Zootaxa 4743(4): 585–593. DOI: 10.11646/zootaxa.4743.4.8 Paywall Reference page. [diagnosis: p. 586]

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